Flora of New York/Rosoideae 1

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Fabales
Flora of New York — Rosales : Rosaceae : Rosoideae 1 :
Ulmarieae, Rubeae, Colurieae, Sanguisorbeae

Rosales: Rosaceae: Rosoideae 2
Table of
contents
Genus
index
Protected species index Invasive species index


Rosales introduction[edit | edit source]

The order Rosales is part of the nitrogen-fixing clade in the fabids:
fabids

Zygophyllales

COM clade

Celastrales

Malpighiales

Oxalidales

nitrogen‑fixing clade 

Fabales

Rosales

Cucurbitales

Fagales


Rosaceae introduction[edit | edit source]

The grouping of the Rosaceae (rose family) taxa used here is based on the Angiosperm Phylogeny Website,[1] Eriksson (2003),[2] Longhi (2014),[3] Sun et al. (2018).[4] X. Chen et al. (2020).[5]

The subfamilies Rosoideae and Amygdaloideae are present in New York.

Rosaceae
Dryadoideae

Cercocarpus

Dryas

Rosoideae
Ulmarieae

Filipendula

Rosodeae
Colurieae

Geum (avens)

Waldsteinia (barren strawberry)

Rubeae

Rubus (brambles)

Roseae

Rosa (roses)

Potentilleae
Potentillineae

Argentina (siverweed)

Potentilla (cinquefoil)

Fragariinae

Fragaria (strawberries)

Dasiphora (shrubby cinquefoil)

Drymocallis (tall cinquefoil)

Sibbaldia (three-toothed cinquefoil)

Comarum (marsh cinquefoil)

Agrimonieae
Agrimoniinae

Agrimonia (agrimony)

Sanguisorbineae

Poteridium (American burnet)

Poterium (salad burnet)

Sanguisorba (burnet)

Amygdaloideae
Niellieae

Physocarpus

Spiraeeae

Spiraea

Lyonothamneae

Lyonothamnus floribundus

Amygdaleae

Prunus

Sorbarieae

Sorbaria (false spiraea)

Osmaronieae

Exochorda (pearlbush)

Kerrieae

Rhodotypos (jetbead)

Kerria (Japanese rose)

Gillenieae

Gillenia

Maleae
Lindleyinae

Kageneckia

Vauquelinia

Malinae

Crataegus (hawthorn)

Amelanchier (shadbush)

Cydonia (quince)

Pyracantha (firethorn)

Sorbus (mountain ash)

Pyrus (pear)

Photinia

Cotoneaster

Chaenomeles (quince)

Malus (apple)

Aronia (chokeberry)


Rosoideae introduction[edit | edit source]

The Rosaceae subfamily Rosoideae contains the following tribes:
  • Agrimonieae (Acaena, Agrimonia, Poteridium, Poterium, Sanguisorba)
  • Colurieae (Fallugia, Geum, Sieversia, Waldsteinia)
  • Potentilleae (Alchemilla, Aphanes, Chamaerhodos, Comarum, Dasiphora, Drymocallis, Duchesnea, Fragaria, Horkelia, Horkeliella, Ivesia, Potentilla, Sibbaldia, Sibbaldiopsis)
  • Roseae (Rosa)
  • Rubeae (Rubus)
  • Ulmarieae (Filipendula)

Tribe Ulmarieae[edit | edit source]

Filipendula[edit | edit source]

Filipendula rubra
Filipendula contains about ten Eurasian species and two North American species. Of these, one Midwestern and two Eurasian species have been found to have naturalized in New York.

Filipendula rubra (queen of the prairie) is the only North American native Filipendula found in New York State, where it is only known to be present in the wild as a garden escape.[1] It can be found on roadsides, ditches, thickets, and disturbed soils, but it has been less common than the Eurasian Filipendula ulmaria.[2]



Rosales — Rosaceae — Rosoideae — Ulmarieae — Filipendula
Genus Filipendula Queen, Dropwort N.Y. Status Images Distribution NY NPT
(Hill) B.L.Rob.

1768. Ulmaria rubra  Hill
1770. Spiraea lobata  Gronov. ex Jacq.
1906. Filipendula rubra  B.L.Rob.
1836. Thecanisia angustifolia  Raf.
1836. Thecanisia purpurea  Raf.
1837. Thecanisia lobata  (Gronov.) Raf.
1879. Filipendula lobata  (Gronov.) Maxim.
1891. Spiraea rubra  (Hill) Britton
1924. Filipendula lobata  var. albiflora  Bergmans
1924. Filipendula lobata  var. venusta-magnifica  Bergmans
Queen-of-the-prairie,
Prairie dropwort,
Red queen-of-the-meadow
Filipendule rouge
Introduced from
 US midwest,
N. America native,
Cultivated in NY,
G4-G5

FACW

Perennial,
Herb-forb,
Part shade

NYFA: X
NYFA: 13 counties
25 counties

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(L.) Maxim.

1753. Spiraea ulmaria  L.
1819. Spiraea denudata  J.& C.Presl
1822. Spiraea ulmaria  f. tomentosa  Hayne)
1838. Thecanisia ulmaria  (L.) Raf.
1879. Filipendula ulmaria Maxim.
1879. Filipendula ulmaria  var. denudata  (J.Presl & C.Presl) Maxim.
1889. Filipendula denudata  Fritsch
1889. Spiraea ulmaria  f. tomentosa  (Hayne) St.-Lag. (not validly publ.)
1894. Ulmaria ulmaria  (L.) Barnhart (not validly publ.)
1895. Thecanisia ulmaria  (L.) Raf. ex B.D.Jacks. (comb. not validly publ.)
1909. Filipendula ulmaria  ssp. denudata  Hayek
Queen-of-the-meadow,
Meadowsweet,
Meadow dropwort,
Elm queen-of-the-meadow,
White queen-of-the-meadow
Reine-des-prés
Introduced from
 temperate Eurasia,
Potentially invasive,
 Invasive.org: 4,
Note:[1][2]

FAC-FACW

Perennial,
Herb-forb

NYFA: X
NYFA: 21 counties
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16 counties

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 Moench

1753. Spiraea filipendula  L.
1768. Ulmaria filipendula  (L.) Hill
1782. Filipendula hexapetala  Gilib. (opus utique oppr.)
1794. Filipendula vulgaris Moench
1822. Spiraea vulgaris  (Moench) Gray (nom. superfl.)
1824. Spiraea filipendula  var. vulgaris  Cambess.
1879. Filipendula hexapetala  Gilib. superfl.
1788. Ulmaria filipendula  (L.) Hill ex Focke
1894. Filipendula filipendula  (L.) Voss (not validly publ.)
1915. Ulmaria angustifolia  Rehder
Dropwort,
Common queen-of-the-meadow
Filipendule vulgaire,
Filipendule
Introduced from
 temperate Eurasia,
Not naturalized

NYFA: X
NYFA: Madison, Chemung (1938)
iNat
No RG observations

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Tribe Rubeae[edit | edit source]

The Rubeae include the genera Rubus and Dalibarda (which is here treated as a monospecific subgenus of Rubus.

Rubus[edit | edit source]

Rubus subg. Chamaemorus[edit | edit source]

Rubus chamaemorus
The native status of cloudberry (Rubus chamaemorus) in New York could be subject to question. It is a circumboreal bramble, native to all provinces of Canada and present in the states of Alaska, Minnesota, New Hampshire, and Maine. However, the only vouchered report of cloudberry growing outside of cultivation in New York was from Montauk in Suffolk County, Long Island in 1908.[1][2]
Rosales — Rosaceae — Rosoideae — Rubeae — Rubus subg. Chamaemorus
Genus Rubussubg. Chamaemorus Cloudberry N.Y. Status Images Distribution NY NPT
 L.

1753. Rubus chamaemorus L.
1906. Chamaemorus anglica  Greene
1906. Chamaemorus norwegica  Greene
1920. Chamaemorus chamaemorus (L.) House (not validly publ.)
Cloudberry,
Baked-apple-berry,
Salmon-berry,
Malka
Chicouté,
Ronce petit-mûrier,
Mûres blanches
Native, CoC: 10,
No recent reports,
Impersistent

FACW

Perennial,
Herb-subshrub,
Sun

NYFA: Z
NYFA: Suffolk (1908)
No NY observations

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Rubus subg. Dalibarda[edit | edit source]

Rubus repens
Dalibarda has been treated as a monospecific genus, but the Flora of North America Editorial Committee (2014)[1] treats it as a subgenus of Rubus. Its leaves superficially resemble those of violets, but its flowers and fruit are clearly rubusesque.

There continues to be disagreement about whether R. dalibarda or R. repens should be the accepted species name. Rubus repens was found to be invalid but has been proposed for conservation as the accepted name.[2]

Subgenus Dalibarda does appear to form a strongly-supported clade with the rest of Rubus and may nest within Rubus.[3]


Rosales — Rosaceae — Rosoideae — Rubeae — Rubus subg. Dalibarda
Genus Rubussubg. Dalibarda Dewdrop N.Y. Status Images Distribution NY NPT
(L.) Kuntze

1753. Dalibarda repens  L.
1762. Rubus dalibarda  L. (nom. superfl.)
1770. Rubus repens  Weston (nom. rej. prop.)
1803. Dalibarda violioides  Michx.
1806. Dalibarda cordata  Stephan (nom. superfl.)
1879. Rubus dalibarda  f. caespitosus  Kuntze
1879. Rubus dalibarda  f. laciniostipulatus  Kuntze
1891. Rubus repens  (L.) Kuntze (nom. cons. prop.)
Dewdrop,
Robin runaway,
False-violet,
Star-violet
Dalibarde rampante
Native, CoC: 9,
Likely secure

FAC

Perennial,
Herb-forb,
Part shade - shade

NYFA: 4
iNat

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Rubus subg. Anoplobatus[edit | edit source]

Rubus odoratus
purple-flowering raspberry
The thornless Anoplobatus subgenus contains Rubus odoratus.
Rosales — Rosaceae — Rosoideae — Rubeae — Rubus subg. Anoplobatus
Genus Rubussubg. Anoplobatus Raspberry N.Y. Status Images Distribution NY NPT
 L.

1753. Rubus odoratus L.
1892. Rubus odoratus  var. columbianus  Millsp.
1903. Rubacer odoratum  Rydb.
1903. Rubacer columbianum  Rydb.
1906. Bossekia odorata  Greene
1912. Bossekia columbiana  A.Heller
Purple-flowering raspberry,
Purple-flowered raspberry,
Virginia raspberry,
Thimbleberry
Ronce odorante,
Framboisier sauvage,
Calottes
Native, CoC: 7,
Secure



Perennial,
Subshrub,
Sun - shade

NYFA: 5

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Rubus subg. Idaeobatus[edit | edit source]

Rubus idaeus
red raspberry
The Rubus subgenus Idaeobatus species are generally referred to as raspberries.
Rosales — Rosaceae — Rosoideae — Rubeae — Rubus subg. Idaeobatus
Genus Rubussubg. Idaeobatus Raspberry N.Y. Status Images Distribution NY NPT
 L.
 ssp. strigosus (Michx.) Focke

1803. Rubus strigosus  Michx.
1872. Rubus idaeus  var. strigosus  (Michx.) Maxim.
1906. Batidaea acalyphacea  Greene
1906. Batidaea strigosa  (Michx.) Greene
1907. Rubus egglestonii  Blanch.
1911. Rubus idaeus  ssp. strigosus  (Michx.) Focke
1912. Batidaea egglestonii  (Blanch.) A.Heller
1913. Rubus acalyphaceus  (Greene) Rydb.
1913. Rubus carolinianus  Rydb.
1913. Rubus arizonicus  (Greene) Rydb.
1913. Rubus subarcticus  (Greene) Rydb.
1945. Rubus aliceae  L.H.Bailey
American red raspberry,
North American red raspberry,
Grayleaf red raspberry
Framboisier sauvage
Native, CoC: 3,
Secure,
S5, G5-T5

FACU-FAC

Perennial,
Shrub

NYFA: 5

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 L.
 ssp. idaeus

1753. Rubus idaeus  L.
1770. Rubus cannabifolius  Weston
1770. Rubus elegantissimus  Weston
1770. Rubus idaeus  var. albus  Weston
1770. Rubus idaeus  var. biflorus  Weston
1770. Rubus idaeus  var. laevis  Weston
1789. Rubus daeus ruber  Aiton
1796. Rubus fragrans  Salisb.
1811. Rubus obtusifolius  Willd.
1915. Batidaea idaea  (L.) Nieuwl.
1915. Batidaea vulgaris  Nieuwl.
European red raspberry,
Framboisier
Introduced from
 Eurasia,
SNA, G5-T5

FACU-FAC

Perennial,
Shrub

NYFA: X

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L.

1753. Rubus occidentalis  L.
1794. Rubus tomentosus  Borkh. (nom. superfl.)
1828. Rubus idaeus  var. americanus  Torr.
1839. Rubus occidentalis  var. incisus  Schltdl.
1874. Rubus occidentalis  ssp. euoccidentalis  Focke
1890. Rubus occidentalis  var. japonicus  Miyabe
1891. Rubus occidentalis  var. grandiflorus  Focke
1898. Rubus fruticosus  var. tomentosus  Fiori
1902. Rubus occidentalis  var. pallidus  L.H.Bailey
1905. Rubus occidentalis  var. flavobaccus  Blanch.
1906. Melanobatus occidentalis  (L.) Greene
Black raspberry,
Black cap,
Thimbleberry
Framboisier de Virginie
Native, CoC: 3,
Secure,
S5, G5

Perennial,
Shrub

NYFA: 5

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 Peck (pro sp.)

Rubus occidentalis ×
Rubus idaeus ssp. strigosus

Purple wild raspberry Native,
Not assessed,
SNA, GNA

Perennial,
Vine, Shrub

NYFA: 2?

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 Maxim.

1872. Rubus phoenicolasius  Maxim.
Wineberry,
Japanese wineberry,
Wine raspberry
Introduced from
 temperate Asia,
Highly invasive,
 NYIS: 86%[1]

NYFA: X

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 Focke

1899. Rubus illecebrosus Focke
Strawberry raspberry,
Charming raspberry
Framboisier fraisier
Introduced from
 eastern Asia,
Potentially invasive

NYFA: X

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L.

1753. Rubus parvifolius L.
1784. Rubus triphyllus Thunb.
1833. Rubus purpureus Bunge
Japanese raspberry ...
Small-leaf bramble ...
Trailing raspberry ...

Introduced from
 Asia,
 Australia,
Potentially invasive,
Naturalized,
SNA
NYFA: 1 county
NYFA: Xn
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Rubus subg. Rubus[edit | edit source]

Subgenus Rubus contains the blackberries and dewberries.

Rubus subg. Rubus sect. Allegheniensis[edit | edit source]
Rubus allegheniensis
Rubus L. sect. Allegheniensis contains Rubus allegheniensis, R. alumnus, and R. pennus.[1] Of these, only R. allegheniensis is listed for New York State.
Rosales — Rosaceae — Rosoideae — Rubeae — Rubus subg. Rubus sect. Allegheniensis
Genus Rubussubg. Rubussect. Allegheniensis Blackberry N.Y. Status Images Distribution NY NPT
 Porter

1890. Rubus villosus  var. montanus  Porter
1894. Rubus montanus  (Porter) Porter (later homonym)
1896. Rubus allegheniensis Porter
1906. Rubus pergratus  Blanch.
1906. Rubus amnicola  Blanch.
1945. Rubus avipes  L.H.Bailey
Allegheny blackberry,
Common blackberry,
Highbush blackberry
Ronce des Alléghanys,
Mûrier,
Ronce alléghanienne
Native, CoC: 3,
Secure

FACU

Perennial,
Shrub

NYFA: 5

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Rubus subg. Rubus sect. Canadenses[edit | edit source]
Rubus canadensis
Rubus L. sect. Canadenses contains Rubus canadensis and R. kennedyanus.[1] Of these, only Rubus canadensis is listed for New York State.
Rosales — Rosaceae — Rosoideae — Rubeae — Rubus subg. Rubus sect. Canadenses
Genus Rubussubg. Rubussect. Canadenses Blackberry N.Y. Status Images Distribution NY NPT
 L.

1753. Rubus canadensis  L.
1891. Rubus millspaughii  Britton
1895. Selnorition canadensis  (L.) Raf. ex B.D.Jacks.
1901. Rubus randii  (L.H.Bailey) Rydb.
1931. Rubus kennedyanus  Fernald
1932. Rubus montensis  L.H.Bailey
1944. Rubus acridens  L.H.Bailey
1944. Rubus ulterior  L.H.Bailey
1947. Rubus besseyi  L.H.Bailey
Smooth blackberry,
Smooth bramble,
Thornless blackberry
Native,
Vulnerable

NYFA: 3?

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Rubus subg. Rubus sect. Arguti[edit | edit source]
Rubus pensilvanicus
The Flora of North America Editorial Committee (2014) treats the Yankee blackberry (R. frondosus), Burnham's blackberry (R. burnhamii), and the prickly Florida blackberry (R. argutus) as being among the 72 synonyms of the Pennsylvania blackberry (R. pensilvanicus).[1][2] ITIS also lists Rydberg's blackberry (R. rydbergianus), along with the other three, as a synonym of (R. pensilvanicus).[3] This leaves sect. Arguti equivalent to Rubus pensilvanicus itself.
Rosales — Rosaceae — Rosoideae — Rubeae — Rubus subg. Rubus sect. Arguti
Genus Rubussubg. Rubussect. Arguti Blackberry N.Y. Status Images Distribution NY NPT
Poir.

1804. Rubus pensilvanicus Poir.
1822. Rubus argutus  Link
1824. Rubus frondosus  Bigelow
1913. Rubus brainerdii  Rydb.
1932. Rubus abactus  L.H.Bailey
1933. Rubus penetrans  L.H.Bailey
1945. Rubus barbarus  L.H.Bailey
1945. Rubus bellobatus  L.H.Bailey
1945. Rubus blakei  L.H.Bailey
1945. Rubus burnhamii  L.H.Bailey
1945. Rubus rydbergianus  L.H.Bailey
Pennsylvania blackberry,
Pennsylvania raspberry
Ronce de Pennsylvanie
Native, CoC: 3,
Secure

NYFA: 5
iNat

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Rubus subg. Rubus sect. Cuneifolii[edit | edit source]
Rubus cuneifolius

Rosales — Rosaceae — Rosoideae — Rubeae — Rubus subg. Rubus sect. Cuneifolii
Genus Rubussubg. Rubussect. Cuneifolii Blackberry N.Y. Status Images Distribution NY NPT
Pursh

1788. Rubus parvifolius  Walter (sensu auct.)
1814. Rubus cuneifolius Pursh
1925. Rubus cuneifolius  var. angustior  L.H.Bailey
1932. Rubus audax  L.H.Bailey
1938. Rubus longii  Fernald
1941. Rubus cuneifolius  var. subellipticus  Fernald
1943. Rubus chapmanii  L.H.Bailey
1943. Rubus georgiensis  L.H.Bailey
1943. Rubus humei  L.H.Bailey
1943. Rubus randolphiorum  L.H.Bailey
1943. Rubus probativus  L.H.Bailey
1970. Rubus dixiensis  H.A.Davis, A.M.Fuller & T.Davis
Sand blackberry Native, CoC: 10,
Endangered,
No recent reports

Perennial,
Shrub

NYFA: 1
NYFA: 3 counties
iNat
2 counties

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Rubus subg. Rubus sect. Setosi[edit | edit source]
The Flora of North America Editorial Committee (2014) treats both the Bristly Oswego blackberry (R. dissimilis) and the Vermont blackberry (R. vermontanus) as being among the 40 synonyms of the Bristly blackberry (R. setosus).[1][2]
Rosales — Rosaceae — Rosoideae — Rubeae — Rubus subg. Rubus sect. Setosi
Genus Rubussubg. Rubussect. Setosi Blackberry N.Y. Status Images Distribution NY NPT
Bigelow

1824. Rubus setosus Bigelow
1901. Rubus nigricans  Rydb. (nom. illeg.)
1904. Rubus vermontanus  Blanch.
1904. Rubus vermontanus  var. viridifolius  Blanch.
1906. Rubus groutianus  Blanch.
1907. Rubus semisetosus  Blanch.
1932. Rubus wheeleri  L.H.Bailey
1941. Rubus dissimilis  L.H.Bailey
1941. Rubus lawrencei  L.H.Bailey
1947. Rubus beatus  L.H.Bailey
1947. Rubus condignus  L.H.Bailey
1947. Rubus viridifrons  L.H.Bailey
Bristly blackberry,
Setose blackberry,
Small bristleberry,
Bristly raspberry
Ronce sétuleuse
Native, CoC: 2,
Likely secure,
S4

FACW

Perennial,
Shrub

NYFA: 4
iNat

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Rubus subg. Rubus sect. Rubus[edit | edit source]
Rubus laciniatus
Sect. Rubus contains somewhat invasive Old World species of Rubus subg. Rubus. The primary species of this section in New York State is Rubus laciniatus (cutleaf blackberry).

The rare-in-NY but possibly naturalizing and invasive Himalayan blackberry (Rubus bifrons and/or Rubus armeniacus), listed in the New York Flora Atlas only in Bronx and Staten Island (2008), are treated as a single species (R. bifrons) by the Flora of North America. However in Eurasia, they appear to be two distinct species. iNaturalist allows both species, following Kew's POWO, and both have a significant number of research-grade observations in North America.


Rosales — Rosaceae — Rosoideae — Rubeae — Rubus subg. Rubus sect. Rubus
Genus Rubussubg. Rubussect. Rubussubsect. Hiemales Blackberry N.Y. Status Images Distribution NY NPT
Willd.

1806. Rubus laciniatus Willd.
1822. Rubus corylifolius  var. laciniatus  (Willd.) Wallr.
1858. Rubus nemoralis  P.J.Müll. (POWO accepted name 2023)
1893. Rubus vulgaris  var. laciniatus  (Weston) Dippel
1902. Rubus bifrons  var. laciniatus  (Willd.) Murr
1909. Rubus bifrons  var. laciniatus  (Weston) Dalla Torre & Sarnth.
1914. Rubus vulgaris  f. laciniatus  (Weston) Frid.
2014. Rubus nemoralis  f. laciniatus  (Willd.) A.Beek
Cutleaf blackberry,
Cut-leaved blackberry,
Evergreen blackberry,
Cutleaf bramble
Ronce laciniée
Introduced,
Moderately invasive,
 NYIS: 63%[1]

NYFA: X
iNat

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 Vest ex Tratt.

1821. Rubus bifrons Vest
1823. Rubus bifrons  Vest ex Tratt.
1825. Rubus linkianus  Ser.
1846. Rubus nemorosus  var. bifrons  (Vest) Bab.
1874. Rubus armeniacus  Focke (per FNA 2023)
 auct. Rubus procerus  non P.J.Müll. ex Genev
1898. Rubus fruticosus  var. bifrons  (Vest) Fiori
Himalayan berry ...
Himalayan blackberry ...
Himalayan giant blackberry ...
Twice-leaved blackberry ...

Ronce discolore
Introduced from
 Europe,
Moderately invasive,
 NYIS: 57%[2],
No NY specimens

NYFA: X
NYFA: 2 counties
iNat
4 counties

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 Focke
 = Rubus bifrons Vest (per FNA 2023)

1874. Rubus armeniacus Focke
1906. Rubus macrostemon  f. armeniacus  (Focke) Sprib.
1914. Rubus hedycarpus  var. armeniacus  Focke
 auct. Rubus discolor  non Weihe & Nees (1824)
Armenian Blackberry,
Himalayan blackberry,
Himalaya giant blackberry
Introduced from
 Caucasus,
Moderately invasive,
 NYIS: 57%[3],
IO

NYFA: 0
NYFA: Synonymized
1 county

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Schott

1788. Rubus inermis  Pourr.
1809. Rubus inermis  Willd.
1818. Rubus ulmifolius Schott
1824. Rubus discolor  Weihe & Nees
1825. Rubus flagellaris  var. inermis  Ser.
1861. Rubus rusticanus  Mercier
1869. Rubus bellidiflorus  hort. ex K.Koch
1903. Rubus ulmifolius  var. genuinus  Sudre
1903. Rubus ulmifolius  ssp. vulgatus  Sudre
1905. Rubus ulmifolius  var. bellidiflorus  C. K. Schneid.
1914. Rubus ulmifolius  var. inermis  (Ser.) Focke
Elmleaf blackberry ...
Elm-leaved blackberry ...
Introduced,
Potentially invasive

NYFA: Xm
NYFA: Bronx (2016)
iNat
No RG observations

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Rubus subg. Rubus sect. Hispidi[edit | edit source]
Rubus hispidus

Rosales — Rosaceae — Rosoideae — Rubeae — Rubus subg. Rubus sect. Hispidi
Genus Rubussubg. Rubussect. Hispidi Dewberry N.Y. Status Images Distribution NY NPT
 L.

1753. Rubus hispidus  L.
1923. Rubus hispidus  var. blanchardianus  L.H.Bailey
1824. Rubus sempervirens  Bigelow
1940. Rubus hispidus  var. obovalis  Fernald
1941. Rubus blanchardianus  (L.H.Bailey) L.H.Bailey
1941. Rubus fassettii  L.H.Bailey
1941. Rubus hispidus  var. cupulifer  L.H.Bailey
1941. Rubus hispidus  var. pervarius  L.H.Bailey
Bristly dewberry,
Running blackberry,
Swamp blackberry,
Swamp dewberry
Ronce hispide
Native,
Secure

NYFA: 5

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Rubus subg. Rubus sect. Flagellares[edit | edit source]
Rubus flagellaris
The Flora of North America Editorial Committee (2014) treats both the Arundel blackberry (R. arundelanus), Enslen's blackberry (R. enslenii), and the Ithaca blackberry (R. ithacanus) as being among the 112 synonyms of the American dewberry (R. flagellaris).[1][2]
Rosales — Rosaceae — Rosoideae — Rubeae — Rubus subg. Rubus sect. Flagellares
Genus Rubussubg. Rubussect. Flagellares Dewberry N.Y. Status Images Distribution NY NPT
Willd.

1809. Rubus flagellaris Willd.
1894. Rubus baileyanus  Britton
1901. Rubus procumbens  Rydb
1906. Rubus arenicola  Blanch.
1906. Rubus geophilus  Blanch
American dewberry,
Northern dewberry,
Whiplash dewberry,
Northern blackberry
Ronce à flagelles
Native,
Secure

NYFA: 5

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Rubus subg. Rubus sect. Caesii[edit | edit source]
Rubus caesius
Eurasian dewberry
The European dewberry has only been reported outside of cultivation in limited number of counties in North America, including only a few from New York. As of July 2015, it was not listed on New York State's Non-Native Plant Species Invasiveness Assessment.[1] However the Finger Lakes Native Plant Society,[2] Cornell Plantations, and Tompkins County[3] all list Rubus caesius as an invasive plant.
Rosales — Rosaceae — Rosoideae — Rubeae — Rubus subg. Rubus sect. Caesii
Genus Rubussubg. Rubussect. Caesii Dewberry N.Y. Status Images Distribution NY NPT
 L.

1753. Rubus caesius  L.
1782. Rubus caeruleus  Gilib.
1829. Rubus polymorphus  var. caesius  (L.) Spenn.
1888. Rubus caesius intermedia  Schmidely
1889. Rubus caesius  f. eucaesius  Bouvet (not validly publ.)
1891. Rubus caesius turkestanicus  Regel
1894. Rubus fruticosus  ssp. caesius  Bonnier & Layens
1895. Selnorition cesius  (L.) Raf. ex B.D.Jacks.
1898. Rubus fruticosus  var. caesius  (L.) Fiori
1937. Rubus psilophyllus  Nevski (nom. illeg.)
Eurasian dewberry,
European dewberry
Ronce bleue,
Ronce bleuâtre
Introduced from
 Eurasia,
 northern Africa,
Moderately invasive,
CP-3[1]

NYFA: X
NYFA: 5 counties
11 counties

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  1. Appendix 3. Plant species that are considered moderately to highly invasive to natural areas in the central Finger Lakes region. Policy on the use of non-native plants in Cornell Botanic Gardens' accessioned collections (2018)

Rubus subg. Cylactis[edit | edit source]

Rubus pubescens
dwarf raspberry

Rosales — Rosaceae — Rosoideae — Rubeae — Rubus subg. Cylactis
Genus Rubussubg. Cylactis Blackberry N.Y. Status Images Distribution NY NPT
Raf.

1803. Rubus saxatilis  Michx. (nom. illeg.)
1811. Rubus pubescens  Raf.
1819. Cylactis montana  Raf.
1823. Rubus triflorus  Richardson
1824. Rubus canadensis  Torr. (sensu auct.)
1825. Rubus mucronatus  Ser.
1893. Cylactis egopodioides  (Ser.) Raf. ex B.D.Jacks.
1985. Cylactis pubescens  (Raf.) W.A.Weber
Dwarf raspberry,
Dwarf red blackberry,
Creeping red raspberry,
Trailing red raspberry,
Plumboy
Ronce pubescente,
Catherinette
Native, CoC: 8,
Secure,
S5

FACW

Perennial,
Shrub

NYFA: 5

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Tribe Colurieae[edit | edit source]

Appalachian barren-strawberry
The New York members of the Colurieae tribe include about ten species of the genus Geum (avens) and a single species of Waldsteinia (barren-strawberry). Sources are mixed as to whether Waldsteinia should be placed under the Geum genus itself. In either case, the "barren strawberries" of this tribe are more closely related to the Geums than to the other New York strawberries (Fragaria) or mock-strawberries (Duchesnea), which are placed in the Potentilleae tribe below.

Protopopova (2022) concluded: Based on trnH-psbA plastid DNA analysis we believe that the taxonomic latter decision of combining Waldsteinia and Taihangia in one genus with Geum species does not reflect the complex structure of the obtained clade."[1]

Protopopova (2023) concluded: "Based on the present phylogenetic structure of Geum, Waldsteinia, and other closely related taxa, we believe that excluding Waldsteinia together with Coluria, Taihangia, and Oncostylus from Geum s.l. is more justified than ‘lumping’ them together into a single polymorphic genus."[2]


  1. M V Protopopova et al 2021 IOP Conf. Ser.: Earth Environ. Sci. 908 012027 DOI 10.1088/1755-1315/908/1/012027
  2. Protopopova, Marina, Vasiliy Pavlichenko, Victor Chepinoga, Alexander Gnutikov, and Renat Adelshin. 2023. "Waldsteinia within Geum s.l. (Rosaceae): Main Aspects of Phylogeny and Speciation History" Diversity 15, no. 4: 479. https://doi.org/10.3390/d15040479

Waldsteinia[edit | edit source]

Waldsteinia fragarioides
The type species of the genus Waldsteinia was found to be nested within Geum by J. E. E. Smedmark in 2006.[1]

Alan S. Weakley (2023) lists genus Waldsteinia as a "genus of about 7 species, perennial herbs, of disjunct areas in the Northern Hemisphere. Sometimes included in Geum. Protopopova, Pavlichenko, and Stepanov (2021) and Protopopova et al. (2023) supported its treatment as a genus separate from Geum."[2][3]


Rosales — Rosaceae — Rosoideae — Colurieae — Waldsteinia
Genus Waldsteinia Barren strawberry N.Y. Status Images Distribution NY NPT
(Michx.) Tratt.
 ssp. fragarioides

1803.  Dalibarda fragarioides  Michx. in…
1814.  Rubus fragarioides  (Michx.) Dum.Cours. in…
1823.  Waldsteinia fragarioides  (Michx.) Tratt. in…
1825.  Comaropsis fragarioides  (Michx.) DC. ex Ser. in…
1825.  Dryas trifoliata  Pall. ex Ser. in…
1840.  Bossekia fragarioides  (Michx.) Raf in…
2006.  Geum fragarioides  (Michx.) Smedmark in…
Appalachian barren-strawberry
Northern Barren Strawberry
Dry strawberry

Waldsteinie faux-fraisier
Native, CoC: 7,
Secure,
S5, G5

Perennial,
Herb-forb

NYFA: 5
iNat

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Geum[edit | edit source]

Geum canadense

Rosales — Rosaceae — Rosoideae — Colurieae — Geum
Genus GeumL. Avens N.Y. Status Images Distribution NY NPT
Jacq.

1773. Geum canadense  Jacq.
1775. Geum laciniosum  Murray
1788. Geum carolinianum  Walter
1791. Geum album  J.F.Gmel.
1794. Caryophyllata alba  Moench
1831. Geum glutinosum  Pursh ex Lehm.
1832. Sieversia caroliniana  G.Don
1913. Geum camporum  Rydb
1922. Geum canadense  var. camporum  (Rydb.) Fernald & Weath.
1922. Geum canadense  var. grimesii  Fernald & Weath.
1922. Geum canadense  var. texanum  Fernald & Weath.
1937. Geum canadense  var. brevipes  Fernald
White avens,
Canada avens
Benoîte du Canada,
Benoîte blanche
Native, CoC: 5,
Secure,
S5, G5

FAC-FACU

Perennial,
Herb-forb

NYFA: 5
iNat

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 Jacq.

1781. Geum aleppicum Jacq.
1789. Geum strictum  Aiton (nom. superfl.)
1913. Geum decurrens  Rydb.
1824. Geum ranunculoides  Ser.
1829. Geum rugosum  Desf.
1935. Geum aleppicum  var. strictum  (Aiton) Fernald
1939. Geum strictum  var. decurrens  (Rydb.) Kearney & Peebles
1949. Geum aleppicum  ssp. strictum  (Aiton) R.T.Clausen
Yellow avens,
Aleppo avens,
Common avens
Benoîte d’Alep
Native, CoC: 3,
Secure

NYFA: 5

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 L.

1753. Geum rivale  L.
1771. Caryophyllata rivalis  (L.) Scop.
1790. Anemone decaphilla  Krock.
1790. Anemone dodecaphilla  Krock.
1791. Anemone decapetala  J.F.Gmel.
1840. Bernoullia hybrida  Raf.
1840. Bernoullia nutans  (Raf.) Raf.
1840. Bernoullia rivalis  (L.) Raf.
Purple avens,
Water avens,
Chocolate-root
Benoîte des ruisseaux,
Benoîte pourpre
Native, CoC: 8,
Secure

OBL

NYFA: 5

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Murray

1775. Geum laciniatum  Murray
1935. Geum laciniatum  var. trichocarpum  Fernald
Rough avens,
Floodplain avens
Benoîte lacinicée
Native, CoC: 3,
Secure,
S5, G5

FACW-FAC

Perennial,
Herb-forb

NYFA: 5
iNat

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(Raf.) Torr. & A.Gray

1821. Stylypus vernus  Raf.
1821. Stypostylis vernus  Raf.
1840. Geum vernum  (Raf.) Torr. & A.Gray
Spring avens N. America native,
NY nativity uncertain,
SNR, G5

FACU

Perennial,
Herb-forb

NYFA: 2-3

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 Pursh
 var. triflorum

1814. Geum triflorum  Pursh
1823. Sieversia triflora  (Pursh) R.Br.
1832. Geum pubescens  Hook.
1906. Erythrocoma campanulata  (Greene
1906. Erythrocoma canescens  (Greene
1906. Erythrocoma flavula  (Greene
1906. Erythrocoma triflora  (Pursh) Greene
1936. Geum ciliatum  var. triflorum  (Pursh) Jeps. (nom. illeg.)
1958. Geum canescens  (Greene) Munz
Prairie smoke ...
Old man's whiskers ...
Native, CoC: 10,
Threatened,
S2, G5

FACU

Perennial,
Herb-forb

NYFA: 2

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 L.

1753. Geum virginianum  L.
1785. Caryophyllata virginiana  (L.) Lam.
1813. Geum hirsutum  Muhl. ex Link
1822. Geum heterophyllum  Link (nom. nud.)
1828. Geum virginianum  var. lanceolatum  Raf. (nom. nud.)
1828. Geum virginianum  var. macrophyllum  Raf. (nom. nud.)
1828. Geum virginianum  var. ochroleucum  Raf. (nom. nud.)
1828. Geum virginianum  var. ramosum  Raf. (nom. nud.)
1828. Geum virginianum  var. uniflorum  Raf. (nom. nud.)
1889. Geum album  var. flavum  Porter
1896. Geum flavum  (Porter) E.P.Bicknell
1923. Geum virginianum  var. murrayanum  Fernald
Cream-colored avens ...
Native, CoC: 9,
Threatened,
S2

FACU-FAC

Perennial,
Herb-forb

NYFA: 2

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Willd.
 var. macrophyllum

1809. Geum macrophyllum Willd.
1910. Geum fauriei  H.Lév.
1915. Geum japonicum  var. sachalinense  Koidz.
1924. Geum japonicum  var. fauriei  (H.Lév.) Kudô
1926. Geum sachalinense  (Koidz.) Makino
1952. Geum macrophyllum  var. sachalinense  (Koidz.) H.Hara
1953. Geum aleppicum  var. sachalinense  (Koidz.) Ohwi
1967. Geum perincisum  var. intermedium  B.Boivin
1982. Geum macrophyllum  ssp. fauriei  (H.Lév.) Vorosch.
Largeleaf avens,
Large-leaved avens
Native, CoC: 8,
Endangered,
S1, G5

FACW

Perennial,
Herb-forb

NYFA: 1

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 L.

1753. Geum urbanum  L.
1771. Caryophyllata urbana  (L.) Scop.
1794. Caryophyllata officinalis  Moench
1840. Bernoullia media  Raf.
1893. Geranium pseudomolle  (Pant.) B.D.Jacks.
1974. Geum rivale  ssp. urbanum  (L.) Á.Löve & D.Löve
Herb Bennet, Bennet's root,
Wood avens, Town avens,
Clove-root, Colewort,
St. Benedict's herb
Benoîte commune
Introduced,
Invasive

NYFA: X

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Geum hybrids[edit | edit source]

Geum × intermedium

Rosales — Rosaceae — Rosoideae — Colurieae — Geum
Genus GeumL. Avens N.Y. Status Images Distribution NY NPT
Fr. ex Scheutz

Geum aleppicum ×
Geum rivale

1846. Geum × willdenowii  Fisch. & C.A.Mey. (nom. illeg.)
1870. Geum × aurantiacum Fr. ex Scheutz
1923. Geum × meinshausenii  Gams
Orange avens ...
Hybrid yellow & purple avens ...
Native,
Unranked,
SNA, GNA

Perennial,
Herb-forb

NYFA: X
NYFA: Onondaga (1942)
iNat
no NY observations

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M.Hough

Geum canadense ×
Geum laciniatum

2018. Geum × cortlandicum M.Hough
Cortland avens ...
Hybrid white & rough avens ...
Native,
Unranked,
SNA, GNA

NYFA: N
NYFA: Cortland, Seneca
iNat
Cortland, Seneca

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M.Hough, A.V.Gilman & Chapm.-Lam

Geum aleppicum ×
Geum canadense

2022. Geum × hainesianum M.Hough, A.V.Gilman & Chapm.-Lam
Haines’s avens
Hybrid yellow & white avens ...
Native,
Unranked,
SNA, GNA

NYFA: N
NYFA: Tioga (1942)
iNat
Cortland (cultivated)

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J.-P.Bernard & R.Gauthier

Geum canadense ×
Geum urbanum

1986. Geum × catlingii  J.-P.Bernard & R.Gauthier
Catling's avens ...
Hybrid white & town avens ...

Introduced,
Naturalized,
SNA, GNA

Perennial,
Herb-forb

NYFA: X
NYFA: Cortland (2016)
iNat
6 counties

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Ehrh.

Geum rivale ×
Geum urbanum

1791. Geum × intermedium Ehrh.
1800. Geum urbanum intermedium (Ehrh.) Willd.
1812. Geum urbanum  var. intermedium  (Ehrh.) Mérat
1825. Geum rivale  var. intermedium  (Ehrh.) Ser.
1834. Geum rivale  ssp. intermedium  (Ehrh.) Schübl. & G. Martens
1836. Geum × rivale-urbanum  G.Mey. (not validly publ.)
1841. Geum × willdenowii  J.N.Buek bis
1869. Geum × intermedium  var. willdenowii  (J.N.Buek bis) Asch.
Intermediate avens ...
Hybrid purple & town avens ...
Introduced,
Naturalized,
SNA, GNA

Perennial,
Herb-forb

NYFA: X
NYFA: Cortland (2018)
iNat
Cayuga, Cortland

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J.-P.Bernard & R.Gauthier

Geum laciniatum ×
Geum urbanum

1986. Geum × macneillii J.-P.Bernard & R.Gauthier
McNeill's avens ...
Hybrid rough & town avens ...
Introduced,
Naturalized,
SNA, GNA

Perennial,
Herb-forb

NYFA: X
NYFA: Cortland (2018)
iNat
Cayuga, Cortland

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