Flora of New York/Caryophyllales 3

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A reviewed version of this page, approved on 20 January 2019, was based off this revision.
←
Caryophyllales 2:
Caryophyllaceae 		Flora of New York — Caryophyllales 3:
Amaranthaceae, Aizoaceae, Nyctaginaceae, Phytolaccaceae, Molluginaceae, Montiaceae, Portulacaceae, Cactaceae 		→
Cornales
Table of
contents 		Genus
index 		Protected species index Invasive species index


Family Amaranthaceae

The Amaranthaceae (amaranthus family)

Subfamily Amaranthoideae

The Amaranthoideae

Amaranthus

Amaranthus subg. Acnida

Subgenus Acnida contains the dioecious Amaranthus species.

Caryophyllales — Amaranthaceae — Amaranthoideae — Amaranthus subg. Acnida
Genus Amaranthus subg. Acnida Pigweed N.Y. Status Images Distribution NY NPT
(L.) Sauer
1753. Acnida cannabina L.
    
1955. Amaranthus cannabinus
(L.) Sauer
 Waterhemp pigweed,
Tidal marsh amaranth 		Native, CoC: 4,
Secure

NYFA: 5
USDA-N0
ARS-GRIN
FNA
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(Moq.) J.D.Sauer
1849. Acnida tuberculata Moq.
1955. Amaranthus tuberculatus
(Moq.) J.D.Sauer
1972. Amaranthus rudis J.D.Sauer
 Rough-fruited water-hemp,
Tall water-hemp,
Rough-fruited amaranth 		Native, CoC: 1,
Likely secure
OBL-FACW
Annual,
Herb-forb

NYFA: 4
USDA-NN
ARS-GRIN
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Wikispecies
Amaranthus subg. Amaranthus
[1]
  1. ↑ Dr. Mark Schonbeck, Virginia Association for Biological Farming (2015). "Redroot Pigweed (Amaranthus retroflexus), Smooth Pigweed (A. hybridus), and Powell Amaranth (A. powellii)"
Caryophyllales — Amaranthaceae — Amaranthoideae — Amaranthus subg. Amaranthus
Genus Amaranthus subg. Amaranthus Amaranth N.Y. Status Images Distribution NY NPT
L. (1753)
 Common amaranth,
Redroot amaranth,
Redroot pigweed,
Rough pigweed,
Wild-beet amaranth 		Native,
Secure,
NE-3[1]

NYFA: 5
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Spleen amaranth,
Green pigweed,
Smooth pigweed,
Smooth amaranth,
Hybrid amaranth,
Slim amaranth 		Native,
Questionable,
NE-3[1]

NYFA: X
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Powell’s amaranth,
Green amaranth,
Powell’s smooth amaranth 		Introduced from
 southwest,
 South America,
N. America native

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 L.
 Love-lies-bleeding,
Purple amaranth,
Foxtail amaranth 		Introduced

NYFA: X
USDA-XX
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Wikispecies
 L.
    Amaranthus cruentus L.
    A. hybridus cruentus
    A. paniculatus
 Blood amaranth,
Purple amaranth,
Caterpillar amaranth 		Introduced

NYFA: X
USDA-XX
Images
Wikispecies
L. (1753)
 Prince-of-Wales feather,
Prince’s feather,
Prince’s-feather amaranth 		Introduced

NYFA: X
USDA-X
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Wikispecies
 L.
1753. Amaranthus spinosus L.
    
    
 Spiny amaranth,
Thorny amaranth,
Carelessweed 		Introduced from
 neotropics,
Naturalized

NYFA: Xn
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ARS-GRIN
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  1. ↑ a b Category 3 Plants - widespread non-native species - Eastern Region invasive plants, ranked by degree of invasiveness as based on information from States (1998) US Forest Service
Amaranthus subg. Albersia
Caryophyllales — Amaranthaceae — Amaranthoideae — Amaranthus subg. Albersia
Genus Amaranthus subg. Albersia Pigweed N.Y. Status Images Distribution NY NPT
 Raf.
 Seabeach amaranth,
Coast amaranth 		Native, CoC: 10,
Threatened,
NYNHP: 1[1]

NYFA: 2
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Note: FWS
 L.
1759. Amaranthus albus L.
1894. A. graecizansvar. pubescens
Uline & W.L.Bray
1912. Amaranthus pubescens Rydb.
1945. A. albusvar. pubescens Fernald
 auct. Amaranthus graecizans non L.
 Prostrate pigweed,
Tumble pigweed,
Tumbleweed,
White amaranth 		Introduced from
 tropical America

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 S.Watson
1877. Amaranthus blitoides S.Watson
 auct. Amaranthus graecizans non L.
 Prostrate pigweed,
Prostrate amaranth,
Matweed,
Matweed amaranth 		Introduced from
 NA Southwest

NYFA: X
USDA-XX
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 S.Watson
1877. Amaranthus palmeri S.Watson
 Carelessweed,
Palmer’s amaranth 		Introduced from
 NA Southwest

NYFA: X
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 L.
1753. Amaranthus blitum L.
 Purple amaranth,
Livid amaranth
Amarante livide 		Introduced from
 Eurasia,
 Africa

NYFA: X
USDA-XX
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(Lesp. & Thev.) N. Terracc.
 Crispleaf amaranth,
Crisp-leaved amaranth 		Introduced

NYFA: X
USDA-X0
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 L.
 Low amaranth,
Large-fruited amaranth 		Introduced,
Impersistent

NYFA: X
USDA-X0
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Slender amaranth,
Tropical green amaranth,
Green amaranth 		Introduced,
N. America native,
Not naturalized

NYFA: Xm
USDA-N0
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  1. ↑ New York Natural Heritage Program. 2024. Online Conservation Guide for Amaranthus pumilus. Threatened, S2/G2.

Celosia

Caryophyllales — Amaranthaceae — Amaranthoideae — Celosia
Genus Celosia Cockscomb N.Y. Status Images Distribution NY NPT
 L.
    Celosia argentea L.
Silver cockscomb ⓘ
Introduced,
Not naturalized
UPL
Annual,
Herb-forb

NYFA: Xm
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 L.
    Celosia cristata L.
Fasciated cockscomb ⓘ
Introduced,
Not naturalized

NYFA: Xm
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Subfamily Achyranthoideae

The Achyranthoideae

Achyranthes

The Japanese chaff flower, Achyranthes japonica, has been found to be quite invasive where it has naturalized in Kentucky and surrounding states.[1] Although it has not been found to have naturalized in or near New York, Japanese chaff flower has been placed on the DEC's list of prohibited plants.[2]

Caryophyllales — Amaranthaceae — Amaranthoideae — Achyranthes
Genus Achyranthes L. Chaff flower N.Y. Status Images Distribution NY NPT
 (Miq.) Nakai
1865. A. bidentatavar. japonica Miq.
1920. Achyranthes japonica (Miq.) Nakai
 Japanese chaff flower Introduced from
 eastern Asia,
Highly invasive,
 NYIS: 71%[3],
Prohibited[2]

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  1. ↑ Christopher Evans & David D. Taylor (2011) "New Invader Profile: Japanese Chaff Flower - Achyranthes japonica". Wildland Weeds, Summer/Fall 2011.
  2. ↑ a b New York State Department of Environmental Conservation Regulation 6 NYCRR Part 575 Prohibited and Regulated Invasive Species
  3. ↑ New York non-native plant invasiveness assessment – Achyranthes japonica: High (71).

Subfamily Gomphrenoideae

The Gomphrenoideae ...

Alternanthera

Caryophyllales — Amaranthaceae — Gomphrenoideae — Alternanthera
Genus Alternanthera Joyweed N.Y. Status Images Distribution NY NPT
 Kunth
1753. Achyranthes repens L.
1817. Alternanthera pungens Kunth
1821. Alternanthera repens (L.) Link
non J.F.Gmel.
1915. Achyranthes leiantha (Seub.) Standl.
 Khakiweed,
Khaki joyweed 		Introduced,
Imperistent,
No specimens

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Froelichia

Froelichia gracilis (slender snakecotton) is commonly found growing in the disturbed sandy soils around railroads.

Caryophyllales — Amaranthaceae — Gomphrenoideae — Froelichia
Genus Froelichia Snakecotton N.Y. Status Images Distribution NY NPT
 (Hook.) Moq.
1840. Oplotheca gracilis Hook.
1849. Oplotheca texana A.Braun
1849. Froelichia gracilis (Hook.) Moq.
1903. F. texana A.Braun ex Small illeg.
1917. F. braunii Standl.
 Slender cottonweed,
Slender cotton-weed,
Slender snakecotton,
Slender snake-cotton 		Introduced from
 sw North America,
Moderately invasive,
 NYIS: 53%[1]

NYFA: X
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  1. ↑ New York non-native plant invasiveness assessment – Froelichia gracilis: Moderate (53).

Gomphrena

Caryophyllales — Amaranthaceae — Gomphrenoideae — Gomphrena
Genus Gomphrena Globe-amaranth N.Y. Status Images Distribution NY NPT
 L.
1753. Gomphrena globosa L.
    
    
 Globe-amaranth,
Common globe amaranth,
Bachelor's-button 		Introduced,
Impersistent,
Cultivated

NYFA: Xm
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 Gill. ex Moq.
1849. Gomphrena martiana Gill. ex Moq.
    
    
 Globe amaranth Introduced,
Impersistent

NYFA: Xm
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Subfamily Polycnemoideae

The Polycnemoideae ... NYFA includes Polycnemum in the Chenopodiaceae.

Polycnemum

Caryophyllales — Amaranthaceae — Polycnemoideae — Polycnemum
Genus Polycnemum Needleleaf N.Y. Status Images Distribution NY NPT
A.Braun ex Bogenh.
1841. Polycnemum majus A.Braun
    Polycnemum arvense
var. simplex Wallr.
    Polycnemum arvense
ssp. majus Briq.
 Giant needleleaf,
Giant needle-leaf
Grand polycnème 		Introduced from
 Europe,
 Turkey,
Impersistent
Annual,
Herb-forb

NYFA: Xm
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Subfamily Salicornioideae

The Salicornioideae ...

Tribe Salicornieae

Salicornia
Caryophyllales — Amaranthaceae — Salicornioideae — Salicornia
Genus Salicornia Glasswort,
Pickleweed 		N.Y. Status Images Distribution NY NPT
 Standl.
1753. Salicornia virginica L. p.p.
1907. Salicornia europaea
var. pachystachya (W.D.J.Koch) Fernald
1916. Salicornia depressa Standl.
1947. Salicornia europaea
var. simplex (Pursh) Fernald
auct.. Salicornia europaea non L.
 Glasswort,
Virginia glasswort,
American glasswort,
Low glasswort,
Saltwort
Salicorne de Virginie 		Native,
Secure
OBL
Perennial,
Herb-forb,
Subshrub

NYFA: 5
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 Michx.
    Salicornia ambigua
    Salicornia perennis in…
    Salicornia virginica in…
    Sarcocornia pacifica in…
Perennial glasswort ⓘ
Native, CoC: 8,
Likely secure
OBL
Perennial,
Herb-forb

NYFA: N4
Images
Wikispecies
 Torr.
1858. Salicornia bigelovi Torr.
 auct. Salicornia mucronata non Lag.
 Dwarf saltwort,
Dwarf glasswort 		Native,
Threatened

NYFA: 2-3
USDA-
Images
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Sarcocornia
Caryophyllales — Amaranthaceae — Salicornioideae — Sarcocornia
Genus Sarcocornia Swampfire N.Y. Status Images Distribution NY NPT
Samphire,
Chicken claws,
Glasswort,
Saltwort 		Native,
Likely secure

NYFA: 4
Images
Wikispecies

Subfamily Camphorosmoideae

The Camphorosmoideae

Tribe Suaedeae

Suaeda
Caryophyllales — Amaranthaceae — Camphorosmoideae — Suaedeae — Suaeda
Genus Suaeda Forssk. ex J.F. Gmel. 1776 Seepweed N.Y. Status Images Distribution NY NPT
(Hook.) Moq.
1838. Chenopodium calceoliforme
Hook.
1840. Suaeda calceoliformis Moq.
1874. Suaeda depressavar. erecta
1907. Suaeda americana
(Pers.) Fernald
1909. Suaeda erecta A.Nelson
1912. Dondia calceoliformis Rydb.
 Horned sea-blite,
Pursh seepweed,
Pursh’s seepweed,
American sea-blite,
Broom seepweed
Suéda couché 		Native,
Likely secure

NYFA: 4
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 (Elliott) Moq.
1817. Salsola linearis Elliott
1840. Suaeda linearis Moq.
1849. Chenopodina linearis Moq.
1898. Dondia linearis A.Heller
 Annual seepweed,
Narrow-leaf sea-blite,
Southern sea-blite 		Native,
Endangered

NYFA: 1
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Bassett & Crompton
1978. Suaeda rolandii
Bassett & Crompton
 Roland's sea-blite
Suéda de Roland 		Native,
Endangered,
NYNHP: 1[1]

NYFA: 1
USDA-NN
VASCAN
ITIS
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 (L.) Dumort.
1753. Chenopodium maritimum L.
1827. Suaeda maritima Dumort.
 Herbaceous seepweed,
White sea-blite
Suéda maritime 		Introduced from
 North Atlantic coast,
N. America native

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  1. ↑ New York Natural Heritage Program. 2024. Online Conservation Guide for Suaeda rolandii. Endangered, S1/G1-2.

Tribe Camphorosmeae

Bassia
The genus Bassia here includes Kochia scoparia. The Kochia genus was merged into Bassia by A. J. Scott in 1978. Although that merger was not widely accepted as a whole, Kochia scoparia subsp. scoparia has been shown to be embedded within a clade with Bassia hyssopifolia, making the Bassia scoparia name more appropriate for that particular taxon. The same study showed Bassia hirsuta to be in a separate clade, and the new monotypic genus Spirobassia was coined to contain that species.[1]
  1. ↑ Gudrun Kadereit & Helmut Freitag (2011) "Molecular Phylogeny of Camphorosmeae (Camphorosmoideae, Chenopodiaceae): Implications for biogeography, evolution of C4-photosynthesis and taxonomy" TAXON 60(1), February 2011: 51–78
Caryophyllales — Amaranthaceae — Camphorosmoideae — Camphorosmeae
Genus Bassia All. Smotherweed N.Y. Status Images Distribution NY NPT
 Schrad.
1753. Chenopodium scoparium L.
1809. Kochia scoparia Schrad.
1809. Kochia scopariassp. scoparia
1908. Kochia alata Bates
1978. Bassia scoparia A.J.Scott
 Summer-cypress,
Burningbush,
Common kochia,
Mexican fireweed,
Red belvedere
Kochia à balais 		Introduced from
 Eurasia,
Moderately invasive,
 NYIS: 69%[1]

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 (Pall.) Kuntze
1771. Salsola hyssopifolia Pall.
1802. Kochia hyssopifolia Roth
1804. Suaeda hyssopifolia Pall.
1840. Echinopsilon hyssopifolium Moq.
1891. Bassia hyssopifolia Kuntze
 Fivehorn smotherweed,
Five-horned smother-weed,
Five-hooked bassia
Bassia à feuilles d'hysope 		Introduced from
 Eurasia,
Potentially invasive

NYFA: X
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  1. ↑ New York non-native plant invasiveness assessment – Kochia scoparia (L.) Schrad.: 69 .
Spirobassia
Caryophyllales — Amaranthaceae — Camphorosmoideae — Camphorosmeae
Genus Spirobassia Freitag & G.Kadereit Contorted smotherweed N.Y. Status Images Distribution NY NPT
(L.) Freitag & G.Kadereit
1753. Chenopodium hirsutum L.
1834. Echinopsilon hirsutum Moq.
1867. Bassia hirsuta Asch.
2011. Spirobassia hirsuta Freitag & al.
 Hairy smother-weed Introduced

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Subfamily Salsoloideae

The Salsoloideae ...

Tribe Salsoleae

In 2007, Hossein Akhani et al. transferred a number of Old World Salsola species to the resurected genus Kali Mill. (1754).[1] As of 2014, the North American sources used here had not chosen to abide by this transferal (though Wikipedia, Wikispecies, and Wikimedia Commons had made the change several years earlier).
  1. ↑ Hossein Akhani, Gerald Edwards & Eric H. Roalson (2007). "Diversification of the Old World Salsoleae s.l. (Chenopodiaceae): Molecular Phylogenetic Analysis of Nuclear and Chloroplast Data Sets and a Revised Classification". International Journal of Plant Sciences Vol. 168, No. 6 (July/August 2007), pp. 931-956. The University of Chicago Press.
Salsola
Caryophyllales — Amaranthaceae — Salsoloideae — Salsoleae
Genus Salsola L. 1753 Russian thistle N.Y. Status Images Distribution NY NPT
 L.
ssp. kali
1753. Salsola kali L.
1827. Salsola turgida Dumort
2011. Kali turgida (Dumort) Guterm.
[1]
 Russian thistle,
Common saltwort,
Salt-bush 		Introduced from
 Europe

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 L.
ssp. pontica (Pall.) Mosyakin
1803. Salsola kalivar. pontica Pall.
1936. Salsola pontica Iliin
1996. Salsola kalissp. pontica Mosyakin
1999. Salsola tragusssp. pontica Rilke
2010. Kali pontica (Pall.) Sukhor.
[2]
 Russian thistle,
Prickly Russian thistle 		Introduced,
No sepecimens

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 Pall.
1803. Salsola collina Pall.
1913. Salsola chinensis Gand.
1974. Salsola kalissp. collina
Bolòs & Vigo in Butl.
2007. Kali collina Akhani & E.H.Roalson
[3]
 Slender Russian thistle,
Tumbleweed 		Introduced from
 Eurasia

NYFA: X
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 L..
1756. Salsola tragus L.
1771. Kali tragus (L.) Scop.
1881. Salsola kalissp. tragus Nyman
1908. Salsola kalissp. iberica
Sennen & Pau
 Russian thistle,
Tumbleweed,
Prickly Russian thistle
Soude roulante,
Chardon de Russie,
Soude de Ruthénie 		Introduced from
 Eurasia,
 northern Africa

NYFA: X
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  1. ↑ W. Gutermann (2011) "Notulae nomenclaturales 41–45. Neue Namen bei Cruciata und Kali sowie einige kleinere Korrekturen. (New names in Cruciata, Kali, and some small corrections)". Phyton (Horn, Austria) 51 (1): 95–102.
  2. ↑ A. Sukhorukov, J. Akopian & A. Zernov (2010) "On some new and critical taxa of the family Chenopodiaceae in the Flora of the Caucasus"
  3. ↑ Hossein Akhani, Gerald Edwards & Eric H. Roalson (2007). "Diversification of the Old World Salsoleae s.l. (Chenopodiaceae): Molecular Phylogenetic Analysis of Nuclear and Chloroplast Data Sets and a Revised Classification". International Journal of Plant Sciences Vol. 168, No. 6 (July/August 2007), pp. 931-956. The University of Chicago Press.

Subfamily Corispermoideae

The Corispermoideae ...

Corispermum

Caryophyllales — Amaranthaceae — Corispermoideae — Corispermum
Genus Corispermum Bugseed N.Y. Status Images Distribution NY NPT
(Nutt.) Nutt.
var. americanum
1753. C. hyssopifolium auct. non L.
1786. C. orientale auct. non Lam.
1818. C. hyssopifolium  var. americanum  Nutt.
1834. C. americanum Nutt.
1903. C. marginale Rydb
1910. C. simplicissimum Lunell
 American bugseed
Corisperme à feuilles d'hysope 		Introduced,
N. America native

NYFA: X
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 Steven
1753. C. hyssopifolium auct. non L.
1817. C. pallasii Steven
1898. C. hyssopifolium  var. leptopterum  Asch.
1929. C. leptopterum (Asch.) Iljin
1929. C. sibiricum Iljin
 Pallas bugseed,
Siberian bugseed
Corisperme à fruit ailé 		Introduced,
N. America native

NYFA: X
USDA-XN
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Kit. ex Schult.
1814. Corispermum nitidum Kit.
 Shiny bugseed,
Slender bugseed 		Introduced from
 Europe

NYFA: Xm
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ARS-GRIN
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Subfamily Chenopodioideae

The tribal organization used here for the Chenopodioideae is from Kadereit et. al. (2010).[1]

Tribe Axyrideae

Axyris
Caryophyllales — Amaranthaceae — Chenopodioideae — Axyrideae — Axyris
Genus Axyris Russian-pigweed N.Y. Status Images Distribution NY NPT
 L.
1753. Axyris amaranthoides L.
 Russian-pigweed,
Upright axyris
Ansérine de Russie 		Introduced from
 Eurasia,
Invasive

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  1. ↑ Gudrun Kadereit, Evgeny V. Mavrodiev, Elizabeth H. Zacharias, Alexander P. Sukhorukov (2010). Molecular phylogeny of Atripliceae (Chenopodioideae, Chenopodiaceae): Implications for systematics, biogeography, flower and fruit evolution, and the origin of C4 photosynthesis. American Journal of Botany 97(10): 1664–1687.

Tribe Dysphanieae

Dysphania
Caryophyllales — Amaranthaceae — Chenopodioideae — Dysphanieae — Dysphania
Genus Dysphania Glandular-goosefoot N.Y. Status Images Distribution NY NPT
(L.) Mosyakin & Clemants
1753. Chenopodium ambrosioides L.
1985. Teloxys ambrosioides W.A.Weber
2002. Dysphania ambrosioides
Mosyakin & Clemants
 Mexican-tea,
Worm-seed 		Introduced

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(L.) Mosyakin & Clemants
1753. Chenopodium anthelminticum L.
1867. C. ambrosioidesvar. anthelminticum
A.Gray
2002. Dysphania anthelmintica
 Wormseed Introduced

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Jerusalem-oak,
Jerusalem oak goosefoot,
Feather-geranium 		Introduced

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Fetid goosefoot,
Fetid glandular-goosefoot 		Introduced from
 Southwest

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Cut-leaved goosefoot Introduced

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Clammy goosefoot,
Ridged goosefoot,
Small crumbweed 		Introduced

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Teloxys
Caryophyllales — Amaranthaceae — Chenopodioideae — Dysphanieae — Teloxys
Genus Teloxys Wormseed N.Y. Status Images Distribution NY NPT
 (L.) Moq.
1753. Chenopodium aristatum L.
1834. Teloxys aristata Moq.
2002. Dysphania aristata
 Pigweed,
Wormseed 		Introduced,
Impersistent

NYFA: Xm
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Cycloloma
Caryophyllales — Amaranthaceae — Chenopodioideae — Dysphanieae — Cycloloma
Genus Cycloloma Pigweed N.Y. Status Images Distribution NY NPT
(Spreng.) J.M.Coult.
1801. Salsola atriplicifolia Spreng.
1894. Cycloloma atriplicifolium J.M.Coult.
 Winged pigweed,
Tumbling-ringwing 		Introduced

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Tribe Anserineae

Anserineae Dumort., Fl. Belg.: 20. 1827. = Spinacieae Moq., Chenop. Monogr. Enum.: 5, 48. 1840.[1]

Blitum
Caryophyllales — Amaranthaceae — Chenopodioideae — Anserineae — Blitum
Genus Blitum L. 1753 Goosefoot N.Y. Status Images Distribution NY NPT
 Schult.
1822. Blitum nuttallianum Schult.
1891. Monolepis nuttalliana Greene
 Poverty-weed,
Nuttall's poverty-weed
Ansérine hastée 		Introduced,
N. America native,
Impersistent

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 L.
ssp. capitatum
1753. Blitum capitatum L.
1753. Blitum capitatumssp. capitatum
[2]
1857. Chenopodium capitatum Ambrosi p.p.
1857. Chenopodium capitatumvar. capitatum
 Strawberry goosefoot,
Strawberry-blight,
Indian-paint, Indian ink,
Blite goosefoot,
Strawberry-spinach
Chénopode capité 		Introduced,
N. America native

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 (L.) C.A.Mey.
1753. Chenopodium bonus-henricus L.
1766. Atriplex bonus-henricus Crantz
1827. Anserina bonus-henricus Dumort.
1829. Blitum bonus-henricus C.A.Mey.
1834. Agathophytum bonus-henricus Moq.
 Good King Henry,
Allgood, Fat-hen,
Perennial goosefoot,
Wild spinach
Chénopode Bon-Henri,
Ansérine bon-henri,
Bon-Henri 		Introduced from
 Europe

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 L.
1753. Blitum virgatum L.
1794. Morocarpus foliosus Moench
1815. Chenopodium virgatum Thunb.
1864. Chenopodium foliosum Asch.
 Leafy goosefoot Introduced from
 Eurasia,
Impersistent

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  1. ↑ Susy Fuentes-Bazan, Pertti Uotila, Thomas Borsch (2012) "A novel phylogeny-based generic classification for Chenopodium sensu lato, and a tribal rearrangement of Chenopodioideae (Chenopodiaceae)". Willdenowia - Annals of the Botanic Garden and Botanical Museum Berlin-Dahlem, 42(1}, 21 June 2012 , pp. 5-24(20).
  2. ↑ S.L.Mosyakin (2013). "New nomenclatural combinations in Blitum, Oxybasis, Chenopodiastrum, and Lipandra (Chenopodiaceae)." Phytoneuron 2013-56: 1–8. Published 12 August 2013. Publication of Blitum capitatum L. subsp. hastatum (Rydb.) Mosyakin, comb. nov. results in this autonym.
Spinacia
Caryophyllales — Amaranthaceae — Chenopodioideae — Anserineae — Spinacia
Genus Spinacia L. 1753 Spinach N.Y. Status Images Distribution NY NPT
 L.
1753. Spinacia oleracea L.
1753. Spinacea oleracea L.
orth. var.
 Spinach
Épinard 		Introduced from
 Eurasia,
Impersistent

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Tribe Chenopodieae

Fuentes-Bazan et al. (2012) moved a number of species out of Chenopodium to make it monophyletic, calling one of the new segregate genera Chenopodiastrum. The genera that were phylogenically between the Chenopodium and Chenopodiastrum clades were already in the tribe Atripliceae, so they placed Chenopodium and the new Chenopodiastrum in Atripliceae, even though the original Atripliceae would then be surrounded by and outnumbered by the former members of the Chenopodieae. But ARS-GRIN, though recognizing the new Chenopodiastrum, places all of these genera in Chenopodieae and calls Atripliceae a synonym.[1] That's the convention used here for this tribe.
  1. ↑ Family: Chenopodiaceae Vent., nom. cons. - subfam. Chenopodioideae - tribe Chenopodieae USDA, ARS, National Genetic Resources Program. Germplasm Resources Information Network (GRIN). National Germplasm Resources Laboratory, Beltsville, Maryland.
Lipandra
Caryophyllales — Amaranthaceae — Chenopodioideae — Chenopodieae — Lipandra
Genus Lipandra Goosefoot N.Y. Status Images Distribution NY NPT
(L.) S.Fuentes, Uotila & Borsch
var. polysperma
1753. Chenopodium polyspermum L.
2012. Lipandra polysperma S.Fuentes et al.
 Manyseed goosefoot,
Many-seeded goosefoot
Chénopode polysperme,
Chénopode à
  nombreuses graines 		Introduced,
Impersistent

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(L.) S.Fuentes, Uotila & Borsch
var. acutifolia Mosyakin
1805. Chenopodium acutifolium Sm.
1828. C. polyspermumvar. acutifolium Gaudin
2013. L. polyspermavar. acutifolia Mosyakin
 Many-seeded goosefoot,
Acute-leaved goosefoot
Chénopode à feuilles aiguës 		Introduced,
Impersistent

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Oxybasis
Caryophyllales — Amaranthaceae — Chenopodioideae — Chenopodieae — Oxybasis
Genus Oxybasis Goosefoot N.Y. Status Images Distribution NY NPT
(L.) S.Fuentes, Uotila & Borsch
var. rubra
1753. Chenopodium rubrumvar. rubrum
2012. Oxybasis rubravar. rubra
[1]
 Red pigweed,
Red goosefoot,
Coast-blite goosefoot
Chénopode rouge,
Ansérine rouge 		Native,
Threatened

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 (L.) S.Fuentes et al.
ssp. glauca
1753. Chenopodium glaucumvar. glaucum
2012. Oxybasis glaucavar. glauca
[1]
 Oak-leaved goosefoot Introduced,
N. America native

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(L.) S. Fuentes, Uotila & Borsch
1753. Chenopodium urbicum L.
2012. Oxybasis urbica in…
Urban goosefoot ⓘ
City goosefoot ⓘ
Introduced from
 Eurasia,
N. America native

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(L.) S.Fuentes, Uotila & Borsch
1771. Blitum chenopodioides L.
1933. Chenopodium chenopodioides
Aellen
2012. Oxybasis chenopodioides
S.Fuentes et al.
 Low goosefoot,
Saltmarsh goosefoot
Chénopode à feuilles
 grasses 		Introduced from
 Eurasia,
 northern Africa,
 western N. America

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Genus Oxybasis (excluded taxa) Goosefoot N.Y. Status Images Distribution NY NPT
 (L.) S.Fuentes et al.
var. humilis (Hook.) Mosyakin
1838. Chenopodium humile Hook.
1880. C. rubrumvar. humile S.Watson
1971. C. rubrumssp. humile Hultén
2013. O. rubravar. humilis Mosyakin
 Coast-blite goosefoot,
Marshland goosefoot 		Introduced from
 western N. America,
Canada native

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  1. ↑ a b sensu S.L.Mosyakin (2013). "New nomenclatural combinations in Blitum, Oxybasis, Chenopodiastrum, and Lipandra (Chenopodiaceae)." Phytoneuron 2013-56: 1–8. Published 12 August 2013.
Chenopodiastrum
Caryophyllales — Amaranthaceae — Chenopodioideae — Chenopodieae — Chenopodiastrum
Genus Chenopodiastrum S.Fuentes et al. 2012 Goosefoot N.Y. Status Images Distribution NY NPT
(Torr.) S.Fuentes, Uotila & Borsch
1827. Chenopodium hybridumvar. simplex Torr.
1832. Chenopodium simplex Raf.
1929. Chenopodium gigantospermum Aellen
2012. Chenopodiastrum simplex S.Fuentes &al.
 Giant-seeded goosefoot,
Maple-leaved goosefoot 		Native,
Secure

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(L.) S.Fuentes, Uotila & Borsch
1753. Chenopodium murale L.
2012. Chenopodiastrum murale S.Fuentes &al.
 Nettle-leaved goosefoot,
Australian-spinach 		Introduced from
 Eurasia,
 northern Africa

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Atriplex
Caryophyllales — Amaranthaceae — Chenopodioideae — Atripliceae — Atriplex
Genus Atriplex L. 1753 Saltbush N.Y. Status Images Distribution NY NPT
 Humb. & Bonpl.
ex Willd.
1806. A. cristata Humb. & Bonpl. ex Willd.
    A. arenaria Nutt.
    A. littoralis (Jacq.) Fawc. & Rendle
    A. pentandra (Jacq.) Standl.
 Crested saltbush Native,
Vulnerable

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 Raf.
non (Nutt.) J.F.Macbr. (1918) nom. illeg.
1817. Atriplex dioica Raf.
1818. Chenopodium subspicatum Nutt.
1874. A. patulavar. subspicata S.Watson
1906. Atriplex subspicata Rydb.
 Saline saltbush,
Thickleaf orach,
Thick-leaved orach 		Native,
Endangered

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 Edmondston
1845. Atriplex glabriuscula Edmondston
1923. Atriplex patulassp. glabriuscula
H.M. Hall & Clem.
 Northeastern saltbush,
Glabrous orach,
Seaside orach,
Scotland Orache,
Bracted orache
Arroche glabriuscule 		Native,
Endangered,
Note:[1]

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Creeping saltbush,
Fat-hen,
Thin-leaved orach,
Hastate-leaved orache,
Triangle orache 		N. America native

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 L.
1753. Atriplex patula L.
1753. Atriplex litoralis L.
1804. Atriplex angustifolia Sm.
1872. Teutliopsis patula Čelak.
1902. A. hastatassp. patula E.Pons
1904. A. hastatavar. patula Farw.
1923. A. patulassp. typica H.M.Hall & Clem.
 Halberd-leaved saltbush,
Spear orach,
Spearscale
Arroche étalée,
Arroche des champs 		Introduced from
 Eurasia,
 northern Africa

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 L.
1753. Atriplex hortensis L.
1753. Atriplex hortensisvar. rubra L.
1803. Atriplex nitens Schkuhr
 Garden orache,
Garden saltbush,
French spinach,
Mountain-spinach
Arroche des jardins,
Arroche rouge des jardins,
Bonne-dame 		Introduced from
 southern Europe?

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Frosted orache,
Belgian orach 		Introduced

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Tumbling orach,
Tumbling saltweed 		Introduced

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Tartarian saltbush Introduced

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 L.
    
    
    
    
 Grassleaf orache Introduced,
No specimens

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  1. ↑ NYFA is not certain of which of the three varieies of Atriplex glabriuscula is/are found in New York. This may place the status in question.
Chenopodium
The native and naturalized Chenopodium (gooosefoot) species found in New York are all members of subgenus Chenopodium, section Chenopodium, as described in Flora of North America North of Mexico.[1]
  1. ↑ Steven E. Clemants & Sergei L. Mosyakin (2003) "38.7. Chenopodium - Goosefoot." Flora of North America North of Mexico. Vol. 4.
Chenopodium subsect. Favosa
Caryophyllales — Amaranthaceae — Chenopodioideae — Chenopodieae — Chenopodium
Genus Chenopodium subsect. Favosa Goosefoot N.Y. Status Images Distribution NY NPT
 Moq.
var. bushianum (Aellen) Cronquist
1929. Chenopodium bushianum Aellen
????. C. berlandierivar. acutidentatum Aellen
1991. C. berlandierivar. bushianum Cronquist
 auct. C. paganum non Reichenb.
 Bush's goosefoot,
Bush's pigweed,
Pit-seeded goosefoot 		Native,
Secure

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 Moq.
var. boscianum Wahl
1840. Chenopodium boscianum Moq
1954. C. berlandierivar. boscianum Wahl
 Pitseed goosefoot,
Pit-seeded goosefoot 		Native,
Endangered

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 Moq.
var. macrocalycium Cronquist
C. macrocalycium Aellen
 Pitseed goosefoot,
Pit-seeded goosefoot,
Large-calyx Goosefoot 		Native,
Endangered

NYFA: 1-2
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Genus Chenopodium subsect. Favosa (excluded taxa) Goosefoot N.Y. Status Images Distribution NY NPT
 Standl.
1916. Chenopodium neomexicanum Standl.
1916. Chenopodium arizonicum Standl.
1916. Chenopodium palmeri Standl.
 New Mexico goosefoot Introduced,
US West native

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Chenopodium subsect. Standleyana
Caryophyllales — Amaranthaceae — Chenopodioideae — Chenopodieae — Chenopodium
Genus Chenopodium subsect. Standleyana Goosefoot N.Y. Status Images Distribution NY NPT
 Aellen
 Standley's goosefoot Native,
Threatened

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Chenopodium subsect. Chenopodium
Caryophyllales — Amaranthaceae — Chenopodioideae — Chenopodieae — Chenopodium
Genus Chenopodium subsect. Chenopodium Goosefoot N.Y. Status Images Distribution NY NPT
 L.
1753. Chenopodium album L.
1820. C. browneanum Roem. & Schult.
1825. C. giganteum D. Don
 Lambs-quarters,
White goosefoot,
Pigweed
Ansérine blanche 		Introduced,
N. America native

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Schrad. ex W.D.J.Koch & Ziz
1775. Chenopodium triangulare Forssk.
1814. Chenopodium opulifolium Schrad.
1962. C. albumssp. opulifolium Maire
 Seaport goosefoot,
Grey goosefoot 		Introduced from
 Eurasia,
 Africa

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 Roth
    
 Striped goosefoot N. America native,
Unk. NY nativity

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 L.
1753. Chenopodium vulvaria L.
 Stinking goosefoot,
Arrach,
Vulvaria 		Introduced,
Impersistant,
Not naturalized

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Chenopodium subsect. Leptophylla
Caryophyllales — Amaranthaceae — Chenopodioideae — Chenopodieae — Chenopodium
Genus Chenopodium subsect. Leptophylla Goosefoot N.Y. Status Images Distribution NY NPT
 Rydb.
 Desert goosefoot,
Narrow-leaved goosefoot 		Introduced,
N. America native

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Genus Chenopodium subsect. Leptophylla (excluded taxa) Goosefoot N.Y. Status Images Distribution NY NPT
    
    
    
    
 Aridland goosefoot,
Narrow-leaved goosefoot 		Introduced

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Chenopodium subsect. Fremontia
Caryophyllales — Amaranthaceae — Chenopodioideae — Chenopodieae — Chenopodium
Genus Chenopodium subsect. Fremontia (excluded taxa) Goosefoot N.Y. Status Images Distribution NY NPT
    
    
    
    
 Fremont's goosefoot N. America native
 western,
N.Y. excluded 		File:Budd's flora of the Canadian Prairie Provinces (1987) (20232160418).jpg

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Subfamily Betoideae

The Betoideae

Caryophyllales — Amaranthaceae — Betoideae
Genus Beta Beet N.Y. Status Images Distribution NY NPT
 L.
1753. Beta vulgaris L.
1762. Beta maritima L.
1882. Beta vulgarisssp. maritima Arcang.
 Beet,
Common beet
Betterave 		Introduced from
 Eurasia,
 northern Africa

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Family Aizoaceae

The Aizoaceae (the fig-marigold family or ice plant family) ...

Subfamily Tetragonioideae

Tetragonia

Caryophyllales — Aizoaceae — Tetragonioideae
Genus Tetragonia Tetragonia N.Y. Status Images Distribution NY NPT
(Pallas) Kuntze
 New Zealand spinach Introduced,
Impersistent

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Subfamily Sesuvioideae

Sesuvium

Caryophyllales — Aizoaceae — Sesuvioideae
Genus Sesuvium Sea purslane N.Y. Status Images Distribution NY NPT
 (Walter)
Britton, Sterns & Poggenb.
 Slender sea-purslane,
Puerto Rico sea-purslane 		Native,
Endangered

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Family Nyctaginaceae

The Nyctaginaceae (four-o'clock family)

Tribe Nyctagineae

Mirabilis
Caryophyllales — Nyctaginaceae — Nyctagineae — Mirabilis
Genus Mirabilis Four-o'clock N.Y. Status Images Distribution NY NPT
Heart-leaved umbrella wort ⓘ
Heartleaf four o'clock ⓘ
Introduced,
N. America native

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(Walter) Heimerl
1893. Mirabilis hirsuta (Pursh) MacMill.
    Mirabilis albida (Walter) Heimerl
Pale umbrella wort ⓘ
White four-o'clock ⓘ
Introduced,
N. America native 		File:Botanical and vegetation survey of Carter County, Montana, Bureau of Land Management-administered lands (1998) (20206300854).jpg

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var. linearis
 Narrow-leaved four o'clock,
Narrowleaf four o'clock 		Introduced,
N. America native,
Unk. naturalization

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 L.
Marvel-of-Peru ⓘ
Marvel of Peru ⓘ
Introduced

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Family Phytolaccaceae

The Phytolaccaceae (pokeweed family)...[1]

Phytolacca

Phytolacca americana
American pokeweed
  1. ↑ Phytolaccaceae Troy Weldy & David Werier (2013) New York Flora Atlas. New York Flora Association, Albany, New York.
Caryophyllales — Phytolaccaceae — Phytolacca
Genus Phytolacca Pokeweed N.Y. Status Images Distribution NY NPT
 L.
var. americana
1753. Phytolacca americana L.
1762. Phytolacca decandra L.
 American pokeweed,
Inkberry,
Pigeonberry,
Pokeberry,
Virginia poke
Raisin d'Amérique 		Native,
Secure
Perennial,
Herb-forb

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Family Molluginaceae

Mollugo verticillata
carpet-weed
The Molluginaceae (family) ... [1]
  1. ↑ Molluginaceae Troy Weldy & David Werier (2013) New York Flora Atlas. New York Flora Association, Albany, New York.

Mollugo

Caryophyllales — Molluginaceae
Genus Mollugo Carpetweed N.Y. Status Images Distribution NY NPT
 L. (1753)
 Carpet-weed,
Green carpetweed,
Indian chickweed,
Mollugo verticillé 		Introduced,
N. America native

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Family Montiaceae

The Montiaceae Raf. (1820) was adoped by APG III (2009) to house a number of genera sgregated from the Portulacaceae (purslane family). Montiaceae includes the spring ephemerals in the genus Claytonia.

Claytonia

Claytonia caroliniana
Carolina spring-beauty
Claytonia contains the two New York spring ephemerals C. caroliniana (Carolina springbeauty) and C. virginica (eastern spring beauty). These two species can be most readily be differentiated by the shape of their leaves. C. virginica is generally found in drier and thinner soils than C. caroliniana.
Claytonia leaf shape leaf length leaf width
C. caroliniana broad oblanceolate to oval 25-75 mm 10-30 mm
C. virginica linear to narrow oblanceolate 75-180 mm 2-15 mm
Caryophyllales — Montiaceae — Claytonia
Genus Claytonia Spring beauty N.Y. Status Images Distribution NY NPT
 Michx.
1803. Claytonia caroliniana Michx.
1807. Claytonia spathulifolia Salisb.
1966. C. carolinianavar. spathulifolia W.H.Lewis
1975. C. carolinianavar. lewisii McNeill
 Carolina springbeauty,
Carolina spring-beauty,
Wide-leaved spring beauty
Claytonie de Caroline,
Claytonie feuille-large 		Native,
Secure

NYFA: 5
USDA-NN
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VASCAN
ITIS
FNA
Tropicos
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Wikispecies
 L.
var. virginica
1753. Claytonia virginica L.
1931. Claytonia robusta (Somes) Rydb.
1831. Claytonia media (DC.) Link
1992. C. virginicavar. hammondiae J.J.Doyle &al.
 Eastern spring beauty,
Virginia springbeauty,
Narrow-leaf spring-beauty,
Narrow-leaved spring beauty
Claytonie de Virginie,
Claytonie feuille-étroite 		Native,
Secure

NYFA: 5
USDA-NN
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VASCAN
ARS-GRIN
ITIS
FNA
Tropicos
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Family Portulacaceae

The Portulacaceae (purslane family) now contains only the genus Portulaca.[1]
  1. ↑ Portulacaceae Troy Weldy & David Werier (2013) New York Flora Atlas. New York Flora Association, Albany, New York.

Portulaca

Portulaca oleracea
common purslane
The origin of Portulaca oleracea is uncertain, but purslane seeds and the pollen of both purslane and corn were discovered in sediments at the bottom of meromitic Crawford Lake, a few miles west of Lake Ontario, indicating that both plants were present there prior to European settlement. It is thought that pre-Columbian people pulled the purslane from their corn fields and then washed it in the lake prior to cooking it.[1] Globally, P. oleracea is widely-naturalized and treated as a noxious weed in many locations.
  1. ↑ R. Byrne; J.H. McAndrews (1975). "Pre-Columbian puslane (Portulaca oleracea L.) in the New World" (PDF). Nature. 253 (5494): 726–727. doi:10.1038/253726a0. Retrieved 27 October 2018.
Caryophyllales — Portulacaceae — Portulaca
Genus Portulaca Purslane N.Y. Status Images Distribution NY NPT
 L.
1753. Portulaca oleracea L.
 Common purslane,
Little hogweed 		Introduced,
Canada native,
Naturalized

NYFA:
USDA-XN
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ARS-GRIN
Images
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 Hook.
 Garden purslane,
Moss-rose,
Rose moss,
Rose-moss-of-garden 		Introduced,
Not naturalized

NYFA: X
USDA-XW
Images
Wikispecies

Family Cactaceae

The Cactaceae (cactus family) has just one taxon native to New York.[1]
  1. ↑ Cactaceae Troy Weldy & David Werier (2013) New York Flora Atlas. New York Flora Association, Albany, New York.

Subfamily Opuntioideae

Tribe Opuntieae

Opuntia
Opuntia humifusa
eastern prickly-pear
The eastern prickly-pear (Opuntia humifusa) is found only in the southeastern part of New York State and is limited to rocky and sandy areas.
Caryophyllales — Cactaceae — Opuntioideae — Opuntieae — Opuntiea
Genus Opuntia Pricklypear N.Y. Status Images Distribution NY NPT
(Raf.) Raf. (1830)
 Eastern prickly-pear Native,
Likely secure

NYFA: 4
USDA-NN
ARS-GRIN
ITIS
FNA
Images
Wikispecies
←
Caryophyllales 2:
Caryophyllaceae 		Flora of New York — Caryophyllales 3:
Amaranthaceae, Aizoaceae, Nyctaginaceae, Phytolaccaceae, Molluginaceae, Montiaceae, Portulacaceae, Cactaceae 		→
Cornales
Table of
contents 		Genus
index 		Protected species index Invasive species index
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