Flora of New York/Lycophytes

From Wikibooks, open books for an open world
Jump to navigation Jump to search

Dipsacales
Flora of New York — Lycophytes
Pre-polypod
ferns
Table of
contents
Genus
index
Protected species index Invasive species index


Class Lycopodiopsida contains the lycophytes (clubmosses, firmosses and quillworts).[1][2]
tracheophytes
lycophytes
Lycopodiales : Lycopodiaceae
Lycopodielloideae

Lycopodiella spp. (5 bog cubmosses)

Pseudolycopodiella caroliniana (slender bog clubmoss)

Lycopodioideae

Dendrolycopodium spp. (3 tree cubmosses)

Diphasiastrum spp. (8 ground cedars)

Spinulum spp. (3 bristly clubmosses)

Lycopodium spp. (2 running clubmosses)

Huperzioideae

Huperzia spp. (4 firmosses)

Isoetales : Isoetaceae

Isoetes spp. (8 quillworts)

Selaginellales : Selaginellaceae

Selaginella spp. (3 spikemosses)

euphyllophytes

Polypodiopsida (ferns)

spermatophytes (seed plants)


Class Order Family Subfamily Genus Common names #
Lycopodiopsida

(lycophytes)
Lycopodiales Lycopodiaceae Lycopodielloideae Lycopodiella bog clubmoss 5
Pseudolycopodiella false bog clubmoss 1
Lycopodioideae Lycopodium clubmoss 2
Dendrolycopodium tree clubmoss 3
Spinulum interrupted clubmoss 3
Diphasiastrum ground cedar 7
Huperzioideae Huperzia firmoss 5
Isoetales Isoetaceae Isoetes quillwort 8
Selaginellales Selaginellaceae Selaginella spikemoss 3

Order Lycopodiales[edit | edit source]

The Lycopodiales ...

Family Lycopodiaceae[edit | edit source]

The Lycopodiaceae (clubmoss family) can, for identification purposes in New York State, roughly be divided into clumping (firmoss) and creeping (clubmoss) groups.
Lycopodiaceae
Huperzioideae (firmoss)

Huperzia lucidula (shining firmoss)

Huperzia appressa (mountain firmoss)

Huperzia selago (northern firmoss)

Huperzia × bartleyi (Bartley’s firmoss)

Lycopodielloideae
bog clubmoss

Lycopodiella inundata (northern bog-clubmoss)

Lycopodiella appressa (southern bog-clubmoss)

Lycopodiella alopecuroides (foxtail bog-clubmoss)

slender bog-clubmoss

Pseudolycopodiella caroliniana (Carolina clubmoss)

Lycopodioideae
running clubmoss

Lycopodium clavatum (staghorn clubmoss)

Lycopodium lagopus (one-cone clubmoss)

tree clubmoss

Dendrolycopodium obscurum (flat-branched tree clubmoss)

Dendrolycopodium dendroideum (prickly tree clubmoss)

Dendrolycopodium hickeyi (Hickey’s tree clubmoss)

bristly clubmoss

Spinulum annotinum (common bristly clubmoss)

Spinulum canadense (northern bristly clubmoss)

groundcedar

Diphasiastrum digitatum (southern ground cedar)

Diphasiastrum tristachyum (blue ground cedar)

Diphasiastrum complanatum (northern ground cedar)

Diphasiastrum sitchense (Sitka ground cedar)

clumping
(firmoss)
creeping
(clubmoss)

Subfamily Huperzioideae[edit | edit source]

The Huperzioideae, known as firmosses, have also been treated as a separate family Huperziaceae. Firmosses differ from other Lycopodiaceae in that they grow in clumps, instead of producing creeping, horizontal stems like other family members.[1] Of the three genera in this group, Huperzia is a temperate and arctic genus and is the only one found naturally in New York. Of the other two genera, Phlegmariurus is native to tropical and subtropical regions, and Phylloglossum is native to Australia and New Zealand.
Huperzia[edit | edit source]
Huperzia lucidula, shining firmoss
Commonly known as firmoss or fir-clubmoss, Huperzia (hoo-per-zee-uh) was named for Johann Peter Huperz, an early 19th century German botanist who specialized in ferns.

The rare Appalachian firmoss was published as Huperzia appalachiana in 1992, but authorities, including Weakly (2023)[1] consider it to be indistinct from the previously published Huperzia appressa.


Lycopodiales — Lycopodiaceae — Huperzioideae — Huperzia
Genus HuperziaBernh. (1801) Firmoss N.Y. Status Images Distribution NY NPT
(Michx.) Trevis.

1803.  Lycopodium lucidulum  Michx.
1875.  Huperzia lucidula  (Michx.) Trevis.
1903.  Huperzia selago  ssp. lucidulum  Clute
1937.  Lycopodium lucidulum  var. verum  Wherry
1939.  Urostachys lucidulus  (Michx.) Herter ex Nessel
1939.  Urostachys lucidulus  var. reclinatus  (Dup) Nessel
1965.  Huperzia selago  ssp. lucidula  (Michx.) Á.Löve & D.Löve
1967.  Lycopodium lucidulum  var. tryonii  Mohlenbr.
1999.  Huperzia lucidula  var. tryonii  (Mohlenbr.) Mohlenbr.
Shining firmoss,
Shining clubmoss,
Shining fir-clubmoss
Lycopode brillant,
Huperzie brillant
Native, CoC: 5,
Secure,
S5, G5

FAC-FACW

Perennial,
Subshrub

NYFA: 5
iNat

USDA-NN
POWO
FSUS
Go Botany
VASCAN
ARS-GRIN
ITIS
FNAFNA
Tropicos
NatureServe
IPN
Images
Wikispecies
(Bach.Pyl. ex Desv.) Á.Löve & D.Löve

1827.  Lycopodium selago  var. appressum  Bach.Pyl. ex Desv.
1902.  Lycopodium selago  f. appressum  (Bach.Pyl. ex Desv.) Gelert
1930.  Lycopodium appressum  (Bach.Pyl. ex Desv.) Petrov in…
1939.  Urostachys selago  var. appressus  (Bach.Pyl. ex Desv.) Herter ex Nessel  in…
1956.  Lycopodium selago  ssp. appressa  (Bach.Pyl. ex Desv.) D.Löve
1961.  Huperzia appressa  (Bach.Pyl. ex Desv.) Á.Löve & D.Löve in…
1968.  Lycopodium selago  ssp. appressum  (Bach.Pyl. ex Desv.) Hultén
1981.  Huperzia selago  var. appressa  (Bach.Pyl. ex Desv.) Ching
1992.  Huperzia appalachiana Beitel & Mickel in…
Appalachian firmoss,
Appalachian clubmoss,
Appalachian fir-clubmoss,
Mountain firmoss,
Mountain fir-clubmoss
Lycopode des Appalaches
Native, CoC: 9,
Rare,
S3, G4-G5,
NYNHP: 3[1]

FACU

Perennial,
Subshrub

NYFA: 3
NYFA: Essex, Greene
iNat
Essex County

USDA-NN
COL
GBIF
WFO
POWO
FSUS
Go Botany
VASCAN
ITIS
FNAFNA
NatureServe
IPN
Images
Wikispecies
(L.) Bernh. ex Schrank & Mart.

1753.  Lycopodium selago  L.
1779.  Lycopodium densum  Lam. (nom. superfl.)
1805.  Plananthus selago  (L.) P.Beauv.
1819.  Plananthus fastigiatus  Opiz
1821.  Lycopodium abietiforme  Gray
1829.  Huperzia selago  (L.) Bernh. ex Schrank & Mart.
1831.  Lycopodium selago  var. densum  Wallr.
1866.  Selago vulgaris  Schur
1923.  Urostachys selago  (L.) Herter
1942.  Lycopodium selago  var. typicum  Wherry (not validly publ.)
1955.  Mirmau selago  H.P.Fuchs
1997.  Huperzia acuminata  (Sugim.) Satou
Northern fir-moss,
Fir clubmoss
Lycopode sélagine,
Huperzie sélagine
Native, CoC: 10,
Endangered,
NYNHP: 1[2]

NYFA: 1
NYFA: 5 counties
iNat
2 counties

USDA-NN
FSUS
Go Botany
VASCAN
ARS-GRIN
ITIS
FNA
Tropicos
NatureServe
IPN
Images
Wikispecies
(F.E.Lloyd & Underw.) Holub

1900.  Lycopodium porophilum  F.E.Lloyd & Underw.
1903.  Lycopodium selago  var. porophilum  Clute
1905.  Lycopodium lucidulum  var. porophilum  (F.E.Lloyd & Underw.) Clute
1908.  Lycopodium lucidulum  f. porophilum  (F.E.Lloyd & Underw.) Clute
1923.  Urostachys porophilus  (F.E.Lloyd & Underw.) Herter
1939.  Urostachys lucidulus  var. porophilus  (F.E.Lloyd & Underw.) Nessel
1965.  Huperzia selago  var. porophila  (F.E.Lloyd & Underw.) Á.Löve & D.Löve
1985.  Huperzia porophila  (F.E.Lloyd & Underw.) Holub
Rock clubmoss,
Rock firmoss,
Lloyd's clubmoss
Lycopode porophile
N. America native,
N.Y. excluded

NYFA-Excluded
NYFA: Excluded
iNat
Carbon County, PA

USDA-NN
POWO
FSUS
VASCAN
FNA
Tropicos
Images
Wikispecies
(Clute) Kartesz & Gandhi

1903.  Lycopodium lucidulum  f. occidentale  Clute
1903.  Lycopodium lucidulum  f. occidentalis  Clute
1932.  Lycopodium lucidulum  var. occidentale  (Clute) L.R.Wilson
1966.  Lycopodium selago  f. occidentale  (Clute) B.Boivin
1966.  Lycopodium selago  var. occidentale  (Clute) B.Boivin
1967.  Lycopodium lucidulum  var. tryonii  Mohlenbr.
1991.  Huperzia occidentalis  (Clute) Kartesz & Gandhi
1992.  Huperzia occidentalis  (Clute) Beitel (nom. illeg.)
Pacific Clubmoss ...
Western Clubmoss ...

Lycopode de l'Ouest ...
N. America native
 Northwest,
N.Y. excluded

Perennial,
Herb

NYFA-Excluded
NYFA: Excluded
iNat
Northwest N.A. only

USDA-NN
GBIF
POWO
ITIS
FNA
Tropicos
NatureServe
BONAP
Images
Wikispecies
Huperzia hybrids[edit | edit source]

Lycopodiales — Lycopodiaceae — Huperzioideae — Huperzia
Genus HuperziaBernh. (1801) Firmoss N.Y. Status Images Distribution NY NPT
(Cusick) Holub

Huperzia lucidula ×
Huperzia porophila

1987.  Lycopodium × bartleyi  Cusick
1991.  Huperzia × bartleyi  (Cusick) Holub
1991.  Huperzia × bartleyi  (Cusick) Kartesz & Gandhi
Bartley's clubmoss Native,
Unranked

NYFA: U
NYFA: Ulster County
iNat
No NY obsevations

USDA-N0
POWO
ITIS
Tropicos
Images
Wikispecies
(Abbe) Kartesz & Gandhi

Huperzia lucidula ×
Huperzia selago

1953.  Lycopodium × buttersii  Abbe
1991.  Huperzia × buttersii  Kartesz & Gandhi
Butters' club-moss,
Butters' firmoss
Lycopode de Butters
N. America native,
N.Y. excluded

NYFA-Excluded
iNat

USDA-NN
POWO
Go Botany
VASCAN
ITIS
FNA
Tropicos
NatureServe
Images
Wikispecies
A.Haines

Huperzia appressa ×
Huperzia lucidula

2003.  Huperzia × protoporophila  A.Haines
Hybrid rock firmoss N. America native,
No N.Y. reports

NYFA: D
iNat

Go Botany
VASCAN
Tropicos
IPN
Images
Wikispecies
A.Haines

Huperzia appressa ×
Huperzia selago

2003.  Huperzia × beitelii  A.Haines
2003.  Huperzia × josephbeitelii  A.Haines
Beitel's firmoss,
Joseph Beitel's clubmoss
N. America native,
No N.Y. reports

NYFA: D
iNat

USDA-NN
Go Botany
VASCAN
Tropicos
IPN
Images
Wikispecies

Subfamily Lycopodioideae[edit | edit source]

Lycopodium[edit | edit source]
Running clubmoss
The genus name Lycopodium (lī-kə-ˈpō-dē-əm) was derived from the Greek lukos (wolf) and podion (small foot). It is commonly known as running clubmoss.

Also treated as Lycopodium sect. Lycopodium.


Lycopodiales — Lycopodiaceae — Lycopodioideae — Lycopodium
Genus Lycopodium Running clubmoss N.Y. Status Images Distribution NY NPT
L.

1753.  Lycopodium clavatum  L.
1805.  Lepidotis clavata  (L.) P.Beauv.
1805.  Lepidotis ciliata  P.Beauv.
1805.  Lepidotis inflexa  P.Beauv.
1810.  Lepidotis aristatum  Humb. & Bonpl. ex Willd.
1925.  Lycopodium clavatum  var. laurentianum  Vict.
1925.  Lycopodium clavatum  var. subremotum  Vict.
Common clubmoss,
Running clubmoss,
Running pine,
Running cedar,
Staghorn clubmoss,
Elk moss
Lycopode à massue
Native, CoC: 5,
Secure

NYFA: 5
iNat

USDA-NN
POWO
Go Botany
ARS-GRIN
ITIS
FNAFNA
Images
Wikispecies
(Laest. ex Hartm.) Zinserl. ex Kuzen

1831.  Lycopodium clavatum  var. monostachyon  Hook. & Grev.
1838.  Lycopodium clavatum  var. integrifolium  (Hook. ex Goldie) Spring
1842.  Lycopodium clavatum  var. integerrimum  Spring
1858.  Lycopodium clavatum  var. lagopus  Laest. ex C.Hartm.
1901.  Lycopodium clavatum  var. brevispicatum  Peck
1910.  Lycopodium clavatum  var. megastachyon  Fernald & Bissell
1925.  Lycopodium clavatum  f. furcatum  Vict.
1950.  Lycopodium clavatum  ssp. monostachyon  (Hook. & Grev.) Selander
1953.  Lycopodium lagopus  (Laest. ex C.Hartm.) Zinserl. ex Kuzen
1958.  Lycopodium clavatum  ssp. megastachyon  (Fernald & Bissell) Á.Löve & D.Löve
One-cone clubmoss,
One-cone ground-pine,
Running pine,
Running cedar,
Arctic stag's-horn clubmoss
Lycopode patte-de-lapin
Native, CoC: 4,
Likely secure

NYFA: 4
iNat

USDA-NN
POWO
Go Botany
ITIS
FNAFNA
Images
Wikispecies
Dendrolycopodium[edit | edit source]
Tree clubmoss
From Greek δενδρος (denros: tree), λυκος (lukos: wolf), and ποδιων (podion: legs), Dendrolycopodium (den-drō-lī-kə-ˈpō-dē-əm) contains tree-clubmoss species, which are fairly erect and have branched stems. This genus, also treated as Lycopodium sect. Obscura, is characterized by stalked terminal strobili, which contain their sporangia.
Lycopodiales — Lycopodiaceae — Lycopodioideae — Dendrolycopodium
Genus Dendrolycopodium Tree clubmoss N.Y. Status Images Distribution NY NPT
(L.) A.Haines

1753.  Lycopodium obscurum  L.
1833.  Lycopodium dendroideum  var. obscurum  (L.) Torr. ex Beck
1937.  Lycopodium obscurum  var. genuinum  Wherry
2003.  Dendrolycopodium obscurum  (L.) A.Haines
Ground pine,
Princess-pine,
Eastern tree clubmoss,
Rare clubmoss,
Flat-branch tree clubmoss
Lycopode obscur
Native, CoC: 5,
Secure

FACU

NYFA: 5
iNat

USDA-NN
Go Botany
ARS-GRIN
ITIS
FNAFNA
Images
Wikispecies
(Michx.) A.Haines

1803.  Lycopodium dendroideum  Michx.
1890.  Lycopodium obscurum  var. dendroideum  D.C.Eaton
1916.  Lycopodium obscurum  var. hybridum  Farw.
2003.  Dendrolycopodium dendroideum  (Michx.) A. Haines
Prickly tree clubmoss,
Northern tree clubmoss,
Treelike clubmoss,
Prickle tree club-moss
Lycopode dendroïde
Native, CoC: 5,
Secure

FACU

NYFA: 5
iNat

USDA-NN
Go Botany
ARS-GRIN
ITIS
FNAFNA
Images
Wikispecies
(W.H.Wagner, Beitel, & Moran) A.Haines

1977.  Lycopodium obscurum  var. isophyllum  Hickey
1989.  Lycopodium hickeyi  W.H.Wagner, Beitel & R.C.Moran
2003.  Dendrolycopodium hickeyi  (W.H.Wagner, Beitel, & Moran) A.Haines
Pennsylvania clubmoss,
Hickey's tree club-moss
Native, CoC: 5,
Secure

Perennial,
Herb

NYFA: 5
iNat

USDA-NN
Go Botany
ITIS
FNAFNA
Images
Wikispecies
Spinulum[edit | edit source]
Spinulum annotinum
Spinulum (ˈspīn-yə-ləm) is also treated as Lycopodium sect. Annotina.
Lycopodiales — Lycopodiaceae — Lycopodioideae — Spinulum
Genus Spinulum Bristly clubmoss N.Y. Status Images Distribution NY NPT
(L.) A.Haines

1753. Lycopodium annotinum  L.
1772. Lycopodium dubium  Zoëga
2003. Spinulum annotinum  (L.) A.Haines
Common bristly clubmoss,
Common stiff clubmoss,
Interrupted clubmoss
Lycopode interrompu
Native, CoC: 5,
Secure

NYFA: 5
iNat

USDA-NN
Go Botany
VASCAN
ARS-GRIN
ITIS
FNA
Images
Wikispecies
(Nessel) A.Haines

1772. Lycopodium dubium auct. non Zoëga
1834. Lycopodium pungens  (Desv. ex Spring) Bach.Pyl. ex Iljin
1842. Lycopodium annotinum  var. pungens  Desv.
1842. Lycopodium pungens  La Pylaie ex Desv.
1843. Lycopodium annotinum  var. montanum  Tuck.
1939. Lycopodium canadense  Romhild ex Nessel
1968. Lycopodium annotinum  ssp. pungens  Hultén
1969. Lycopodium annotinum  ssp. dubium  (Zoëga) Kallio, Laine & Y.Mäkinen
2003. Spinulum canadense  (Romhild ex Nessel) A.Haines
Northern bristly clubmoss,
Northern stiff clubmoss,
Northern interrupted clubmoss
Native, CoC: 10,
Endangered

NYFA: 1-2
iNat

Go Botany
VASCAN
Tropicos
Images
Wikispecies

Spinulum annotinum ×
Spinulum canadense

Hybrid bristly clubmoss Native,
Unranked

NYFA: U

Images
Wikispecies
Diphasiastrum[edit | edit source]
Ground cedar (genus Diphasiastrum) is also treated as Lycopodium sect. Complanata.


Lycopodiales — Lycopodiaceae — Lycopodioideae — Diphasiastrum
Genus DiphasiastrumHolub (1975) Ground-cedar N.Y. Status Images Distribution NY NPT
(Dill. ex A.Braun) Holub

1848.  Lycopodium digitatum  Dill. ex A.Braun
1901.  Lycopodium complanatum  var. flabelliforme  Fernald
1911.  Lycopodium flabelliforme  (Fernald) Blanch.
1965.  Diphasium complanatum  ssp. flabelliforme   in…
1975.  Diphasiastrum digitatum (Dill. ex A.Braun) Holub
Southern groundcedar,
Southern ground-cedar,
Fan club-moss,
Southern running-pine
Lycopode en éventail,
Courants verts,
Lycopode aplati variété en éventail
Native, CoC: 4,
Secure,
S5, G5

NYFA: 5
iNat

USDA-NN
Go Botany
VASCAN
ARS-GRIN
ITIS
FNAFNA
Tropicos
IPN
Images
Wikispecies
(Pursh) Holub

1814.  Lycopodium tristachyum  Pursh in…
1840.  Lycopodium clavatum  var. tristachyum  (Pursh) Hook.  in…
1848.  Lycopodium complanatum  ssp. tristachyum  (Pursh) Dostál  in…
1859.  Lycopodium complanatum  ssp. chamaecyparissus   in…
1905.  Lycopodium complanatum  var. tristachyon  Clute  in…
1944.  Diphasium tristachyum  (Pursh) Rothm. in…
1961.  Diphasium chamaecyparissus in…
1975.  Diphasiastrum tristachyum (Pursh) Holub in…
Blue ground-cedar,
Deep-root clubmoss
Native, CoC: 4,
Secure,
S5, G5

NYFA: 5
iNat

USDA-NN
POWO
Go Botany
ITIS
FNAFNA
Tropicos
Images
Wikispecies
(L.) Holub

1753.  Lycopodium complanatum  L. in…
1805.  Lepidotis complanata  (L.) P.Beauv. in…
1840.  Lycopodium anceps  Wallr. in…
1944.  Diphasium complanatum  (L.) Rothm. in…
1958.  Diphasium anceps  (Asch.) Á.Löve & D.Löve in…
1975.  Diphasiastrum complanatum (L.) Holub in…
Northern groundcedar,
Trailing clubmoss,
Northern running-pine,
Lycopode aplati
Native, CoC: 8,
Endangered,
S2, G5

NYFA: 1
iNat

USDA-NN
Go Botany
ARS-GRIN
ITIS
FNAFNA
Images
Wikispecies
(Rupr.) Holub (1975)

1845.  Lycopodium sitchense  Rupr. in…
1916.  Lycopodium complanatum  var. sitchense   in…
1923.  Lycopodium × sabinifolium  var. sitchense   in…
1965.  Lycopodium × sabinifolium  ssp. sitchense   in…
1965.  Diphasium sitchense  (Rupr.) Á.Löve & D.Löve in…
1975.  Diphasiastrum sitchense  (Rupr.) Holub in…
Sitka groundcedar,
Alaskan clubmoss,
Sitka club-moss,
Lycopode de Sitka
Native, CoC: 10,
Endangered,
S1, G5

NYFA: 1
iNat

USDA-NN
Go Botany
ARS-GRIN
ITIS
FNA
Images
Wikispecies
(Willd.) Holub (1975)

Diphasiastrum sitchense ×
Diphasiastrum tristachyum

    Lycopodium × sabinifolium Willd.
    Lycopodium sabinifolium Willd.
    Lycopodium armatum Desv.
Ground-fir,
Savinleaf groundpine,
Savin-leaved groundpine,
Cypress clubmoss
Native,
Unranked,
SNA, GNA

NYFA: 3
iNat

USDA-NN
ARS-GRIN
ITIS
FNA
Images
Wikispecies
(Rouy) Holub (1975)

Diphasiastrum complanatum ×
Diphasiastrum tristachyum

    Lycopodium × zeilleri
    Diphasium zeilleri (Rouy) Damboldt
Zeiller's running-pine,
Zeiller's clubmoss
Native,
Unranked,
SNA, GNA

NYFA: 2
iNat

USDA-NN
ITIS
FNA
Images
Wikispecies
(House) Holub (1975)

Diphasiastrum digitatum ×
Diphasiastrum tristachyum

    Lycopodium × habereri House
    Lycopodium complanatum  var. habereri
    Lycopodium tristachyum  var. habereri
Haberer's clubmoss Native,
Unranked,
SNA, GNA

NYFA: U
iNat

USDA-NN
ITIS
FNA
Images
Wikispecies

Subfamily Lycopodielloideae[edit | edit source]

Lycopodiella[edit | edit source]

Bog clubmoss. (Lycopodiella in Latin means little wolf's-foot.)

Lycopodiales — Lycopodiaceae — Lycopodielloideae — Lycopodiella
Genus LycopodiellaHolub (1964) Bog clubmoss N.Y. Status Images Distribution NY NPT
(L.) Holub

1753. Lycopodium inundatum L.
1805. Plananthus inundatus (L.) P.Beauv.
1852. Lepidotis inundata (L.) Opiz
1912. Lepidotis inundata (L.) Börner
1964. Lycopodiella inundata (L.) Holub
Northern bog clubmoss,
Marsh clubmoss,
Inundated clubmoss
Lycopodielle inondé
Native, CoC: 8,
Secure

OBL

Perennial,
Subshrub

NYFA: 5

USDA-NN
Go Botany
ARS-GRIN
ITIS
FNA
Tropicos
Images
Wikispecies
(Chapm.) Cranfill

1843. Lycopodium inundatum  var. bigelovii  Tuck.
1878. Lycopodium inundatum  var. appressum  Chapm.
1900. Lycopodium appressum F.E.Lloyd & Underw.
1981. Lycopodiella appressa (Chapm.) Cranfill
1985. Lycopodiella bigelowii (Tuck.) Holub
Southern bog clubmoss,
Appressed bog-clubmoss,
Chapman's club-moss
Native, CoC: 7,
Likely secure

FACW

Perennial,
Subshrub

NYFA: 4

USDA-NN
Go Botany
ITIS
FNA
Images
Wikispecies
(L.) Cranfill

1753. Lycopodium alopecuroides L.
1944. Lepidotis alopecuroides (L.) Rothm.
1981. Lycopodiella alopecuroides (L.) Cranfill
Foxtail bog-clubmoss,
Foxtail clubmoss
Native, CoC: 7,
Vulnerable,
S2?, G2

FACW

Perennial,
Subshrub
Kings, Nassau, Richmond, Suffolk
NYFA: 3

USDA-N0
Go Botany
ITIS
FNA
Images
Wikispecies
(Eiger) Cranfill (1981)

L. alopecuroides ×
L. appressa

Copeland's bog clubmoss,
Copeland's clubmoss
Native,
Unranked
Suffolk
NYFA: U

USDA-N0
ITIS
Images
Wikispecies
A.Haines (2003)

L. appressa ×
L. inundata

Gilman's bog clubmoss,
Gilman's clubmoss
Native,
Unranked

NYFA: U

ITIS
Images
Wikispecies
Genus Lycopodiella (excluded taxa) Bog clubmoss N.Y. Status Images Distribution NY NPT
( R.J.Eaton) A.Haines (2002)

L. alopecuroides ×
L. inundata

Lycopodium inundatum var. robustum
Bog clubmoss Native,
N.Y. excluded

NYFA: U

Images
Wikispecies
Pseudolycopodiella[edit | edit source]
Lycopodiales — Lycopodiaceae — Lycopodielloideae — Pseudolycopodiella
Genus Pseudolycopodiella(L.) Holub False bog-clubmoss N.Y. Status Images Distribution NY NPT
(L.) Holub

1753. Lycopodium carolinianum L.
1968. Lycopodiella caroliniana (L.) Pic.Serm.
1983. Pseudolycopodiella caroliniana Holub
1983. Pseudolycopodiella meridionalis Holub
Carolina clubmoss,
Slender clubmoss,
Slender bog club-moss
Native, CoC: 9,
Endangered,
NYNHP: 1[1]

NYFA: 1

USDA-N0
Go Botany
ARS-GRIN
ITIS
FNA
Tropicos
Images
Wikispecies

Order Selaginellales[edit | edit source]

Family Selaginellaceae[edit | edit source]

Selaginella[edit | edit source]

Three spikemoss (Selaginella) species have been found growing wild in New York.
Selaginellales — Selaginellaceae — Selaginella
Genus Selaginella Spikemoss N.Y. Status Images Distribution NY NPT
(L.) Spring

1753.  Lycopodium rupestre  L. in…
1803.  Stachygynandrum rupestre  (L.) P.Beauv. in…
1838.  Selaginella rupestris  (L.) Spring in…
1849.  Selaginella rupestris  var. borealis  Spring  in…
1891.  Lycopodioides rupestris  (L.) Kuntze in…
1900.  Selaginella bourgeaui  Hieron. in…
1908.  Selaginella rupestris  f. borealis  (Spring) Clute  ...
1992.  Bryodesma rupestre  (L.) Soják in…
Rock spikemoss,
Dwarf spikemoss,
Northern selaginella
Sélaginelle des rochers,
Sélaginelle rupestre
Native, CoC: 9,
Likely secure,
S4

UPL

Perennial,
Herb

NYFA-4
iNat

NRCS-NN
POWO
Go Botany
ITIS
FNA
Images
Wikispecies
(L.) Spring

1753. Lycopodium apodum L.
1840. Selaginella apus Spring
1840. Selaginella apoda (L.) Spring
Meadow spikemoss,
Creeping spikemoss
Sélaginelle apode
Native, CoC: 6,
Unranked,
Under review[1]

FACW

NYFA-2?
iNat

NRCS-N0
Go Botany
ITIS
FNA
Images
Wikispecies
W.R.Buck

1977. Selaginella eclipes W.R.Buck
1996. Selaginella apoda  ssp. eclipes  Skoda
Hidden spikemoss,
Buck's meadow spikemoss,
Creeping spikemoss
Native, CoC: 8,
Endangered

FACW

NYFA-1

NRCS-NN
Go Botany
ITIS
FNA
Tropicos
Images
Wikispecies
Genus Selaginella (excluded taxa) Spikemoss N.Y. Status Images Distribution NY NPT
(L.) P. Beauv. ex Mart. & Schrank

1753. Lycopodium selaginoides L.
1805. S. spinosa P.Beauv. illeg.
1841. S. selaginoides (L.) P.Beauv. ex Mart. & Schrank
Club spikemoss ...
Northern spikemoss ...
Northern spike-moss ...
Prickly mountain-moss ...
Lesser-club-moss ...

N. America native,
N.Y. excluded

FACW

Perennial,
Herb

NYFA-Excluded

USDA-NN
Go Botany
ARS-GRIN
Images
Wikispecies

Order Isoetales[edit | edit source]

Family Isoetaceae[edit | edit source]

Isoetes[edit | edit source]

Isoetes echinospora
North American Isoetes lacustris plants have recently been segregated as Isoetes macrospora but can't be easily differentiated except by geography.[1]
Isoetales — Isoetaceae — Isoetes
Genus IsoetesL. Quillwort N.Y. Status Images Distribution NY NPT
A.Braun

1846.  Isoetes engelmannii  A.Braun in…
1846.  Isoetes lacustris  var. microspora  Engelm. in…
1867.  Isoetes engelmannii  var. gracilis  Engelm. in…
1891.  Calamaria engelmannii  (A.Braun) Kuntze in…
1905.  Isoetes engelmannii  var. fontana  A.A.Eaton  in…
1994.  Isoetes × fairbrothersii  J.D.Montgom. & W.C.Taylor in…
Engelmann's quillwort,
Appalachian quillwort
Isoète d'Engelmann
Native, CoC: 6,
Secure,
S5

OBL

Perennial,
Herb, Subshrub,
diploid (2n = 22)

NYFA: 5
NYFA: 23 counties
iNat
Cortland (2019)

USDA-NN
COL
GBIF
WFO
POWO
FSUS
Go Botany
VASCAN
ITIS
FNAFNA
Tropicos
NatureServe
BONAP
IPN
Images
Wikispecies
Durieu
 ssp. muricata A.Löve & D.Löve[1]

1789.  Isoetes setacea  Lam. p.p.
1827.  Isoetes tenella  Léman ex Desv. POWOin…
1864.  Isoetes braunii  Durieu in…
1867.  Isoetes echinospora  var. braunii  Engelm.
1945.  Isoetes muricata  var. braunii  (Engelm.) C.F.Reed  in…
1961.  Isoetes echinospora  ssp. muricata  (Durieu) Á.Löve & D.Löve NYFA
Spiny-spore quillwort,
Spring quillwort,
Braun's quillwort,
Bristlelike quillwort
Isoète à spores épineuses,
Isoète à spores spinuleuses
Native, CoC: 7,
Likely secure,
S4

OBL

Perennial,
Herb,
diploid (2n = 22)

NYFA: 4

USDA-?
Go Botany
VASCAN
ARS-GRIN
ITIS
FNA
Tropicos
Images
Wikispecies
A.Braun ex Engelm.

1867. I. tuckermanii A.Braun ex Engelm.
1900. I. tuckermanii  var. borealis  A.A.Eaton
1981. I. acadiensis Kott
1991. I. tuckermanii  ssp. acadiensis  Taylor & Luebke
Tuckerman's quillwort
Isoète de Tuckerman
Native, CoC: 10,
Likely secure,
tetraploid (2n = 44)

NYFA: 4

USDA-NN
FSUS
Go Botany
ITIS
FNA
Images
Wikispecies
R.Dodge

Isoetes echinospora ×
Isoetes engelmannii

1896. Isoetes eatonii R.Dodge
1900. Isoetes gravesii A.A.Eaton
    Isoetes eatonii  var. gravesii  A.A.Eaton
Eaton's quillwort Native,
Vulnerable

NYFA: 3

USDA-NN
ITIS
Images
Wikispecies
Durieu

1753.  Isoetes lacustris  L. in…
1864.  Isoetes macrospora  Durieu in…
1900.  Isoetes hieroglyphica  A.A.Eaton
1922.  Isoetes macrospora  f. hieroglyphica  N.Pfeiff.  ...
Big-spore quillwort ...
Lake quillwort
Carved quillwort
Warty Quillwort

Isoète lacustre
Isoète hiéroglyphique
Native, CoC: 10,
Rare,
S3

OBL

Perennial,
Herb,
decaploid (2n = 110)

NYFA: 3
NYFA: 5 counties
iNat
St. Lawrence only

USDA-NN
GBIF
Go Botany
ITIS
FNAFNA
Images
Wikispecies
D.F.Brunt.

1846.  Isoetes riparia  var. canadensis  Engelm.  in…
1903.  Isoetes canadensis  var. robbinsii  A.A. Eaton
2015.  Isoetes septentrionalis  D.F. Brunt. in…
Northern shore quillwort,
Northern quillwort
isoète du Nord
Native, CoC: 10,
Endangered

OBL

Perennial,
Herb,
tetraploid (2n = 44)

NYFA: 1

USDA-N0
FSUS
Go Botany
VASCAN
ITIS
FNA
Images
Wikispecies
(Engelm.) D.F.Brunt.

Isoetes engelmannii ×
Isoetes septentrionalis

1898.  Isoetes × dodgei A.A.Eaton
 auct. Isoetes × brittonii
 auct. Isoetes engelmannii × riparia
Hybrid of
 Engelmann's quillwort &
 northern shore quillwort
Native,
No recent reports

NYFA: Z?

USDA-N0
Go Botany
ARS-GRIN
ITIS
Images
Wikispecies
  1. The New York Flora Atlas and New England Wild Flower Society list the spiny-spore quillwort as Isoëtes echinospora ssp. muricata because the leaves of the North American plants of bear stomata, while those of the European plants do not. The other sources used here treat ssp. muricata as a synonym.

Isoetes (excluded)[edit | edit source]

Isoetales — Isoetaceae — Isoetes
Genus Isoetes (excluded taxa) Quillwort N.Y. Status Images Distribution NY NPT
Engelm. ex A.Braun[1]

1846. Isoetes riparia Engelm. ex A.Braun
18??. Isoetes saccharata Engelm. in A.Gray
1891. Calamaria saccharata (Engelm.) Kuntze
1901. Isoetes canadensis (Engelm.) A.A.Eaton
1949. Isoetes riparia  f. saccharata  ( Engelm. ex A.Gray ) Proctor
Shore quillwort,
Canada shore quillwort
Native,
N.Y. excluded

NYFA-Excluded

USDA-NN
Go Botany
ITIS
Tropicos
BONAP
Images
Wikispecies
A.A.Eaton

Isoetes lacustris ×
Isoetes tuckermanii

1900. Isoetes × heterospora A.A.Eaton
    Isoetes tuckermanii  var. harveyi  Clute
Harvey's quillwort Native,
N.Y. excluded

NYFA-Excluded

USDA-NN
Go Botany
ITIS
FNA
Images
Wikispecies
D.F. Brunt. & D.M. Britton

Isoetes septentrionalis ×
Isoetes tuckermanii

New England quillwort Native,
N.Y. excluded

NYFA-Excluded

USDA-N0
Go Botany
ITIS
FNA
Images
Wikispecies
  1. Isoetes riparia was apparently misapplied to reports of Isoetes septentrionalis in New York. (NYFA (2016)