Flora of New York/Boraginales

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Solanales
Flora of New York — Boraginales
Lamiales 1
Table of
contents
Genus
index
Protected species index Invasive species index


Family Boraginaceae[edit]

The Boraginaceae (forget-me-not family)...

Subfamily Boraginoideae[edit]

Tribe Eritrichieae[edit]

Genus Hackelia[edit]
Boraginales — Boraginaceae — Boraginoideae — Eritrichieae — Hackelia
HackeliaOpiz Stickseed N.Y. Status Images Distribution 2
(L.) I.M.Johnst.

1753. Myosotis virginiana L.
1818. Echinospermum virginicum (L.) Lehm.
1819. Rochelia virginiana (L.) Roem.& Schult.
1888. Echinospermum Virginianum
(L.) Britton, Sterns & Poggenb.
1891. Lappula Virginiana (L.) Greene
1891. Cynoglossospermum virginicm
(L.) Kuntze
1923. Hackelia virginiana (L.) I.M.Johnst.
Virginia stickseed,
Beggars lice,
Stickseed
Hackélia de Virginie,
Hackélie de Virginie,
Bardanette de Virginie
N.Y. native,
NYS CoC: 4,
Secure

FACU

Biennial,
Herb-forb
Hackelia virginiana 003-3x4.jpg
Nymap.svg
NYFA-5
USDA-NN
NEW Can

Trop.

LBJ

Images, wsp
 (Wahlenb.) Opiz
ssp. americana
(A. Gray) Á.Löve & D.Löve

1882. Echinospermum deflexum
var. americanum
1891. Lappula deflexa var. americana
1897. Lappula americana Rydb.
1924. H. deflexa var. americana
1938. H. americana Fernald
1975. Lappula deflexa ssp. americana
1982. H. deflexa ssp. americana
Nodding stickseed,
American stickseed
N.Y. native,
NYS CoC: 10,
Endangered

Biennial,
Herb-forb
Hackelia deflexa BB-1913.png
Nymap.svg
NYFA-1
USDA-NN
NEW 

Trop.



Images, wsp
Genus Mertensia[edit]
Boraginales — Boraginaceae — Boraginoideae — Eritrichieae — Mertensia
Mertensia Bluebells N.Y. Status Images Distribution 1
 (L.) Pers. ex Link

1753. Pulmonaria virginica L.
1797. Mertensia pulmonarioides Roth
1829. Mertensia virginica Pers. ex Link
Eastern bluebells,
Virginia bluebells,
Roanoke-bells,
Virginia cowslip
Mertensia de Virginie
N.Y. native,
NYS CoC: 7,
Likely secure

FAC-FACW
Virginia Bluebells - Mertensia virginica, Merrimac Farm Wildlife Management Area, Aden, Virginia - 13722927424.jpg
Mertensia virginica nymap.png
NYFA-4
USDA-NN
NEW 
ARS ITIS
Trop.



Images, wsp
Mertensia (excluded taxa) Bluebells N.Y. Status Images Distribution X
 (Ait.) G. Don.

    Mertensia paniculata (Ait.) G. Don.
Tall bluebells N. America native,
N.Y. excluded
Mertensia paniculata 1012.JPG
Excluded nymap.svg
NYFA-XCLD
USDA-NN



NSE BNA


Images, wsp
(L.) Gray

    Mertensia maritima (L.) Gray
    Pneumaria maritima (L.) Hill
    
Oysterleaf
Oysterplant
Seaside bluebells
Sea bluebells

N. America native,
N.Y. excluded
Mertensia maritima RHu 03.JPG
Excluded nymap.svg
NYFA-XCLD
USDA-NN
NEW 





Images, wsp
Genus Amsinckia[edit]
Boraginales — Boraginaceae — Boraginoideae — Eritrichieae — Amsinckia
Amsinckia Fiddleneck N.Y. Status Images Distribution 2
Fisch. & C.A.Mey.

A. menziesii var. intermedia
Coast fiddleneck,
Common fiddleneck,
Intermediate fiddleneck,
Rancher's fiddleneck,
Coast buckthorn
Introduced,
N. America native,
Impersistent
Amsinckia intermedia NPS-1.jpg
Nymap.svg
NYFA-X
USDA-NN






Images, wsp
 Lehm.

Bugloss fiddle-neck,
Tarweed fiddleneck
Introduced,
N. America native,
Impersistent
Amsinckia lycopsoides 9650.JPG
Nymap.svg
NYFA-X
USDA-NN






Images, wsp
(Lehm.) A. Nelson & J.F.Macbr.

yyyy. Amsinckia menziesii
(Lehm.) A. Nelson & J.F.Macbr.
Small-flowered fiddleneck
Menzies' fiddleneck

Introduced,
N. America native,
Not naturalized
Amsinckia menziesii (5882121934).jpg
Nymap.svg
NYFA-X
USDA-NN






Images, wsp
Genus Lappula[edit]
Boraginales — Boraginaceae — Boraginoideae — Eritrichieae — Lappula
Lappula Stickseed N.Y. Status Images Distribution 2
(Retz.) Dumort.

1753. Myosotis lappula L.
1781. Myosotis squarrosa Retz.
1827. L. squarrosa (Retz.) Dumort.
European stickseed
Bluebur
Bur-forget-me-not
Introduced from
 Europe,
 temperate Asia,
Naturalized

Biennial,
Herb-forb
Lappula squarrosa fruits.JPG
Nymap.svg
NYFA-X
USDA-XX

ARS 




Images, wsp
(S.Watson) Greene

1891. Lappula redowskii
(Hornem.) Greene misapplied
1895. L. redowskii var. occidentalis
1899. L. occidentalis
(S.Watson) Greene
Western stickseed
Flatspine stickseed
Flat-spined stickseed
Flat-spined sheepburr
Introduced,
N. America native,
Not naturalized
Stickseed (Lappula occidentalis) (9474248502).jpg
Nymap.svg
NYFA-X
USDA-NN
NEW 
ARS 
Trop.



Images, wsp
Lappula (excluded taxa) Stickseed N.Y. Status Images Distribution X
(Hornem.) Greene

1813. Myosotis redowskii Hornem.
1891. Lappula redowskii
(Hornem.) Greene
Redowski's stickseed

N. America native,
N.Y. excluded
Lappula redowskii (7281612302).jpg
Excluded nymap.svg
NYFA-XCLD






Images, wsp
Genus Asperugo[edit]
Asperugo is a monotypic genus, containing only Asperugo procumbens, commonly known as madwort.
Boraginales — Boraginaceae — Boraginoideae — Eritrichieae — Asperugo
AsperugoL. Madwort N.Y. Status Images Distribution 1
 L.

Madwort
German madwort
Introduced,
Not naturalized
Asperugo procumbens2.JPG
Nymap.svg
NYFA-X
USDA-XX






Images, wsp

Tribe Boragineae[edit]

Genus Anchusa[edit]
Boraginales — Boraginaceae — Boraginoideae — Boragineae — Anchusa
Anchusa Bugloss N.Y. Status Images Distribution 3
 (L.) M. Bieb.

Lycopsis arvensis
Small alkanet
Small bugloss
Annual bugloss
Introduced

Annual,
Herb-forb
20121018Ochsenzunge Altlussheim3.jpg
Nymap.svg
NYFA-X
USDA-XX






Images, wsp
 Mill.

Anchusa italica
Garden alkanet
Italian bugloss
Introduced Anchusa azurea FlowersCloseup 19April2009 DehesaBoyaldePuertollano.jpg
Nymap.svg
NYFA-X
USDA-XX






Images, wsp
 L.

Common alkanet
Common bugloss
Introduced,
Naturalization unknown
Anchusa officinalis eF.jpg
Nymap.svg
NYFA-X
USDA-XX






Images, wsp
Genus Borago[edit]
Boraginales — Boraginaceae — Boraginoideae — Boragineae — Borago
Borago Borage N.Y. Status Images Distribution 1
 L.

1753. Borago officinalis L.
Borage
Common borage
Introduced from
 Europe,
 western Asia,
 northern Africa,
Unk. naturalization

Annual,
Herb-forb
Borago officinalis in Botanical Garden of Besançon 05.JPG
Nymap.svg
NYFA-X
USDA-XW

ARS 




Images, wsp
Genus Brunnera[edit]
Boraginales — Boraginaceae — Boraginoideae — Boragineae — Brunnera
Brunnera Brunnera N.Y. Status Images Distribution 1
 (Adams) I.Johnst.

Large-leaved brunnera,
Siberian bugloss
Introduced Brunnera macrophylla 01.JPG
Nymap.svg
NYFA-X
USDA-X0






Images, wsp
Genus Pulmonaria[edit]
Boraginales — Boraginaceae — Boraginoideae — Boragineae — Pulmonaria
Pulmonaria Lungwort N.Y. Status Images Distribution 1
 Mill.

Bethlehem lungwort Introduced Pulmonaria saccharata 'Argentea'.jpg
Nymap.svg
NYFA-X
USDA-XX






Images, wsp
Genus Symphytum[edit]
Boraginales — Boraginaceae — Boraginoideae — Boragineae — Symphytum
Symphytum Comfrey N.Y. Status Images Distribution 2
 Lepechin

Prickly comfrey Introduced Symphytum asperum bloemen.jpg
Nymap.svg
NYFA-X
USDA-XX






Images, wsp
 L.

S. uliginosum auct. non Kern.
Common comfrey Introduced SymphytumOfficinale-bloem-roze-hr.jpg
Nymap.svg
NYFA-X
USDA-XX






Images, wsp

Tribe Lithospermeae[edit]

Genus Lithospermum[edit]
Boraginales — Boraginaceae — Boraginoideae — Lithospermeae — Lithospermum
Lithospermum Stoneseed N.Y. Status Images Distribution 6
 Michx.

American gromwell,
American stoneseed
N.Y. native,
Vulnerable
Lithospermum latifolium illustration.svg
Lithospermum latifolium NY-dist-map.png
NYFA-3
USDA-NN






Images, wsp
Weakley, Witsell & D.Estes

1905. Onosmodium hispidissimum Mack.
1972. O. molle ssp. hispidissimum B.Boivin
1995. O. bejariense var. hispidissimum
2011. L. parviflorum Weakley, et al.
0000. L. carolinianumauct. non Lam.
Hairy false gromwell,
Shaggy false gromwell,
Softhair marbleseed,
Soft-hair marbleseed
N.Y. native,
Vulnerable
Onosmodium bejariense var. hispidissimum, Torrey's Flora of NY 2-08.jpg
Lithospermum parviflorum NY-dist-map.png
NYFA-3-4
USDA-NN

ARS 
Trop.



Images, wsp
Note: [1]
 L.

1753. Lithospermum virginianum L.
1846. Onosmodium virginianum A.DC.
1905. O. virginianum var. hirsutum Mack.
Virginia false gromwell,
Wild Job's tears
N.Y. native,
Endangered,
NYNHP: 1[2]
Onosmodium virginianum BB-1913.png
Lithospermum virginianum NY-dist-map.png
NYFA-1
USDA-N

ARS 
Trop.



Images, wsp
Note: [1]
(Walter ex J.F. Gmel.) MacMill.
var. croceum

Carolina puccoon,
Plains puccoon
N.Y. native,
Endangered,
No recent reports
Lithospermum caroliniense.jpg
Lithospermum caroliniense var croceum NY-dist-map.png
NYFA-Z
USDA-NN






Images, wsp
(Michx.) Lehm.

Batschia canescens Michx.
Hoary puccoon N.Y. native,
Likely extirpated
Lithospermum canescens Kaldari 01.jpg
Lithospermum canescens NY-dist-map.png
NYFA-Z
USDA-NN






Images, wsp
 L.

European gromwell,
European stoneseed
Introduced Lithospermum officinale2.JPG
Lithospermum officinale NY-dist-map.png
NYFA-X
USDA-XX






Images, wsp
  1. a b In 2011, because Onosmodium was found to be embedded within Lithospermum, Alan S. Weakley, et. al. transfered the species of Onosmodium to (in some cases back to) Lithospermum.J. Bot. Res. Inst. Texas 5(2): 437 – 455. 2011
  2. New York Natural Heritage Program. 2016. Online Conservation Guide for Lithospermum virginianum. Endangered, S1/Gx.
Genus Myosotis[edit]
Myosotis laxa
smaller forget-me-not
Of the seven forget-me-not (Myosotis L.) species reported in New York State, two of them (M. laxa and M. verna) are native.

The genus name is from the Greek muosōtis meaning "mouse ear", based on the shape of the leaves.

Forget-me-not flowers have 5 petals and 5 sepals, and are generally little more than 1/4-inch in diameter.


Boraginales — Boraginaceae — Boraginoideae — Lithospermeae — Myosotis
MyosotisL. Forget-me-not N.Y. Status Images Distribution 7
 Lehm.

1818. Myosotis laxa Lehm.
Smaller forget-me-not
Bay forget-me-not
Small forget-me-not
Small-flowered forget-me-not
Tufted forget-me-not

Myosotis laxiflore
Myosotis à petite fleur
N.Y. native,
NYS CoC: 3,
Secure

OBL

Biennial,
Herb-forb
Myosotis laxa, 2015-06-12, Mount Lebanon, 01.jpg
Myosotis laxa NY-dist-map.png
NYFA-5
USDA-NN
NEW Can
ARS ITIS


LBJ

Images, wsp
 Nutt.

1818. Myosotis verna Nutt.
1888. Myosotis virginica
(L.) Britton, Sterns & Poggenb.
Spring forget-me-not
Spring scorpion grass

N.Y. native,
NYS CoC: 5,
Secure

FAC-FACU

Annual,
Herb-forb
Myosotis verna BB-1913.png
Myosotis verna NY-dist-map.png
NYFA-5
USDA-NN
NEW Can

Trop.

LBJ

Images, wsp
 L.

1753. Myosotis scorpioides L.
1753. M. scorpioides var. palustris
1756. M. palustris (L.) Nathh.
Water forget-me-not
True forget-me-not
Forget-me-not

Introduced from
 Europe,
 temperate Asia,
Naturalized,
Cultivated

OBL

Perennial,
Herb-forb
MyosotisScorpioides.jpg
Myosotis scorpioides NY-dist-map.png
NYFA-X
USDA-XX
NEW Can
ARS 
Trop.



Images, wsp
 (L.) Hill

1753. M. scorpioides var. arvensis
1764. M. arvensis (L.) Hill
Field forget-me-not,
Rough forget-me-not
Introduced from
 Europe,
 temperate Asia,
 northern Africa,
Naturalized

FACU-UPL

Annual,
Herb-forb
20170512Myosotis arvensis1.jpg
Myosotis arvensis NY-dist-map.png
NYFA-X
USDA-XX
NEW Can
ARS 




Images, wsp
Ehrh. ex Hoffm.

1788. M. scorpioides var. sylvatica
1791. M. sylvatica Hoffm.
1794. M. alpestris F.W.Schmidt
1825. M. oblongata Link
Woodland forget-me-not
Wood forget-me-not
Garden forget-me-not

Introduced from
 Europe,
 temperate Asia,
 tropical Asia,
Naturalized

UPL

Perennial,
Herb-forb
Myosotis sylvatica001.jpg
Myosotis sylvatica NY-dist-map.png
NYFA-X
USDA-XX
NEW Can
ARS 
Trop.

IPN 

Images, wsp
Link ex Roem. & Schult.

1819. Myosotis stricta
Link ex Roem. & Schult.
 auct. Myosotis micrantha
non Pall. ex Lehm.
Small-flowered forget-me-not
Strict forget-me-not
Upright forget-me-not

Introduced from
 Europe,
 northwest Africa

Annual,
Herb-forb
Myosotis stricta W.jpg
Myosotis stricta NY-dist-map.png
NYFA-X
USDA-XX
NEW Can
ARS 




Images, wsp
 Pers.

    Myosotis discolor Pers.
    Myosotis versicolor (Pers.) Sm.
Yellow and blue forget-me-not
Changing forget-me-not

Introduced Changing Forget-me-not 600.jpg
Myosotis discolor NY-dist-map.png
NYFA-X
USDA-XX
NEW Can





Images, wsp
Genus Buglossoides[edit]
Buglossoides arvensis
corn gromwell
Buglossoides is a Eurasian genus with only the single species corn gromwell (Buglossoides arvensis) known to have naturalized in North America, including New York. It is listed as a noxious weed in Washington State.[1]

Other Buglossoides species, including B. purpurocaerulea, B. tenuiflora, and B. zollingeri, have likely been cultivated in North America, but have not been widely reported to have naturalized.[2]


Boraginales — Boraginaceae — Boraginoideae — Lithospermeae — Buglossoides
BuglossoidesMoench (1782) Buglossoides N.Y. Status Images Distribution 1
(L.) I.M.Johnst.

1753. Lithospermum arvense L.
1952. B. arvensis (L.) I.M.Johnst.
Corn gromwell
Corn-gromwell

Introduced from
 Europe,
 Asia,
 northern Africa,
Naturalized
20130501Acker-Steinsame Reilingen6.jpg
Buglossoides arvensis NY-dist-map.png
NYFA-X
USDA-XX
NEW 
ARS ITIS

BNA


Images, wsp

Tribe Cynoglosseae[edit]

Genus Cynoglossum[edit]
Boraginales — Boraginaceae — Boraginoideae — Cynoglosseae — Cynoglossum
Cynoglossum Hound's-tongue N.Y. Status Images Distribution 4
 L.
ssp. boreale (Fernald) A.Haines

    Cynoglossum boreale Fernald
    C. virginianum var. boreale
    C. virginianum ssp. boreale
Northern wild comfrey,
Wild hound's-tongue,
Wild comfrey
N.Y. native,
NYS CoC: 8,
Endangered

Perennial,
Herb-forb
Cynoglossum virginianum var boreale 8-eheep (5097861502).jpg
Nymap.svg
NYFA-1-2
USDA-NN
NEW 
ARS ITIS




Images, wsp
Note: CA[1]
 L.
ssp. virginianum

1753. Cynoglossum virginianum L.
    C. virginianum var. virginianum
    C. virginianum ssp. virginianum
2015. Andersonglossum virginianum
(L.) J.I.Cohen
Southern wild comfrey,
Southern hound's-tongue,
Wild comfrey,
Giant-forget-me-not
N.Y. native,
NYS CoC: 10?,
Endangered

Perennial,
Herb-forb
Cynoglossum virginianum by Todd Crabtree (8497631867).jpg
Nymap.svg
NYFA-Z
USDA-N0
NEW 
ARS ITIS




Images, wsp

1753. Cynoglossum officinale L.
Houndstongue Introduced 20150508Cynoglossum officinale3.jpg
Cynoglossum officinale NY-dist-map.png
NYFA-X






Images, wsp

1906. Cynoglossum amabile
Stapf & J.R. Drumm.
Chinese hound's tongue Introduced Cynoglossum amabile.jpg
Cynoglossum amabile NY-dist-map.png
NYFA-X






Images, wsp

Tribe Echieae[edit]

Genus Echium[edit]
Boraginales — Boraginaceae — Boraginoideae — Echieae — Echium
Echium Viper's bugloss N.Y. Status Images Distribution 2
 L.

Common viper's bugloss,
Blue-weed,
Blue devil
Introduced Echium vulgare Belarus.jpg
Echium vulgare NY-dist-map.png
NYFA-X
USDA-XW






Images, wsp
 L.

Echium lycopsis L. p.p.
Salvation jane,
Paterson's curse
Introduced,
Noxious weed
Echium plantagineum4.JPG
No counties NY-dist-map.png

USDA-X






Images, wsp
 Sm.

Echium vulgare var. pustulatum
Bluedevil,
Blue-weed,
Viper's-bugloss
Introduced,
No specimens
Echium pustulatum 001.jpg
No counties NY-dist-map.png
NYFA-X
USDA-X0






Images, wsp

Subfamily Hydrophylloideae[edit]

Genus Hydrophyllum[edit]

Hydrophyllum contains eight species of waterleaf, all native to North America, including two that are native to New York.

Boraginales — Boraginaceae — Hydrophylloideae — Hydrophyllum
Hydrophyllum Waterleaf N.Y. Status Images Distribution 2
 L.

1753. Hydrophyllum virginianum L.
1942. H. virginianum var. atranthum
Virginia waterleaf,
John's cabbage,
Eastern water-leaf,
Shawnee-salad,
Indian-salad
N.Y. native,
NYS CoC: 6,
Secure

FACW-FAC

Perennial,
Herb-forb
Hydrophyllum virginianum - Virginia Waterleaf.jpg
Nymap.svg
NYFA-5
USDA-NN
NEW 
ARS 
Trop.



Images, wsp
 L.

1759. Hydrophyllum canadense L.
Bluntleaf waterleaf,
Blunt-leaved waterleaf,
Broad-leaf waterleaf,
Canada waterleaf,
Hairy waterleaf,
Maple-leaved waterleaf
Hydrophylle du Canada
N.Y. native,
NYS CoC: 8,
Secure

FACW-FACU

Perennial,
Herb-forb
Hydrophyllum canadense - Bluntleaf Waterleaf 2.jpg
Nymap.svg
NYFA-5
USDA-NN
NEW 
ARS 
Trop.



Images, wsp
Hydrophyllum (excluded taxa) ' N.Y. Status Images Distribution X

    
    
    
Great waterleaf

N. America native
 midwest & south,
N.Y. excluded
Hydrophyllum appendiculatum, 2016-05-16, Beechview, 04.jpg
Excluded nymap.svg
NYFA-XCLD






Images, wsp

Genus Ellisia[edit]

Boraginales — Boraginaceae — Hydrophylloideae — Ellisia
Ellisia Ellisia N.Y. Status Images Distribution 1
 (L.) L.

1753. Ipomoea nyctelea L.
1762. Polemonium nyctelea L.
1763. Ellisia nyctelea L.
1913. Nyctelea nyctelea Britton
Waterpod,
Aunt Lucy,
False babyblueeyes,
Nyctelea
N.Y. native,
NYS CoC: 10,
Likely extirpated

FAC-UPL

Annual,
Herb-forb
Ellisia nyctelea.JPG
Ellisia nyctelea nymap.svg
NYFA-Z
USDA-NN
NEW 
ARS 
Trop.
BNA


Images, wsp

Genus Phacelia[edit]

Boraginales — Boraginaceae — Hydrophylloideae — Phacelia
Phacelia Scorpion-weed N.Y. Status Images Distribution 3
 (L.) Trel. & Small
var. dubia

1753. Polemonium dubium L.
1888. Phacelia dubia Trel. & Small
Small-flowered scorpionweed,
Smallflower phacelia
N.Y. native,
Likely extirpated
Phacelia dubia (L.) Trel. - smallflower phacelia PHDU.png NYFA-Z
USDA-N0


Trop.



Images, wsp
 A.Gray

1878. Phacelia campanularia A.Gray
1925. Phacelia minor var. campanularia Jeps.
Desert scorpion-weed,
Desert bluebells,
California bluebell
Introduced,
from Arizona,
from California
Phacelia-campanularia-20080330.JPG NYFA-X
USDA-NX

ARS 
Trop.



Images, wsp
 Benth.

1835. Phacelia ciliata Benth.
Great valley scorpion-weed,
Great valley phacelia
Introduced,
from California
NYFA-X
USDA-N0


Trop.



Images, wsp

Subfamily Heliotropioideae[edit]

Genus Heliotropium[edit]

Boraginales — Boraginaceae — Heliotropioideae — Heliotropium
Heliotropium Heliotrope N.Y. Status Images Distribution 4
 L.
var. curassavicum

Licorice-weed,
Salt heliotrope,
Seaside heliotrope
Introduced,
US South native,
Impersistent
Heliotropium curassavicum NRCS-1.jpg
Heliotropium curassavicum var curassavicum NY-dist-map.png
NYFA-X
USDA-N0






Images, wsp
 L.

European heliotrope,
Héliotrope d'Europe
Introduced,
Eurasia native,
N. Africa native,
Australia native?
P1000546 Heliotropium europaeum (Boraginaceae) Flower.JPG
Heliotropium europaeum NY-dist-map.png
NYFA-X
USDA-X0






Images, wsp
 L.

Indian heliotrope,
Indian turnsole
Introduced,
Impersistent
Heliotropium indicum W2 IMG 9793.jpg
Heliotropium indicum NY-dist-map.png
NYFA-X
USDA-X0






Images, wsp
 L.

Garden heliotrope Introduced,
Impersistent
1024 Sonnenwende. Heliotropium arborescens-1630.jpg
Heliotropium arborescens NY-dist-map.png
NYFA-X
USDA-X0






Images, wsp

References[edit]