Flora of New York/Polypodiales

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Ferns 1
non-polypod
Flora of New York: Polypodiales
Pinales
Table of
contents
Genus
index
Protected species index Invasive species index


Order Polypodiales[edit]

The order Polypodiales Link (1833) contains about 20 families.

Family Dennstaedtiaceae[edit]

The Dennstaedtiaceae Lotsy (1909), (bracken-fern family)

Dennstaedtiaceae
Dennstaedtia Hay-scented fern N.Y. Status Images Distribution 1
 (Michx.) T.Moore

1803. Nephrodium punctilobulum  Michx.
1810. Aspidium punctilobum  Willd.
1827. Sitobolium punctilobum  Desv.
1848. Dicksonia punctilobula  A.Gray
1857. Dennstaedtia punctilobula  T.Moore
Eastern hay-scented fern,
Hay-scent fern,
Boulder fern

Dennstaedtie à
  lobules ponctués
Native,
Secure
Dennstaedtia punctilobula2.jpg
Dennstaedtia punctilobula nymap.svg
NYFA-5
USDA-NN
NEW Can
ARS ITIS
FNA Trop.


Images, wsp
Pteridium Brackenfern N.Y. Status Images Distribution 2
 (L.) Kuhn
var. latiusculum (Desv.) Underw. ex A. Heller

1827. Pteris latiuscula  Desv.
1909. Pteridium aquilinum var. latiusculum
1914. Pteridium latiusculum  (Desv.) Hieron.
1941. Pteridium aquilinum ssp. latiusculum
Eastern brackenfern,
Eastern bracken,
Brackenfern

Fougère-aigle de l'Est
Native,
Secure
DidzialapisSakys.JPG
Pteridium aquilinum var latiusculum nymap.svg
NYFA-5
USDA-NN
NEW Can
ARS ITIS
FNA Trop.


Images, wsp
 (L.) Kuhn
var. pseudocaudatum (Clute) A. Heller

1900. Pteris aquilina var. pseudocaudata Clute
1900. Pteridium aquilinum var. pseudocaudatum
1919. Pteridium latiusculum var. pseudocaudatum
1941. Pteridium aquilinum ssp. pseudocaudatum
Tailed bracken,
Brackenfern,
Southeastern Bracken,
Western brackenfern
Native,
Endangered,
DEC-A[1]
Bracken frond unfurling - geograph.org.uk - 769807.jpg
Pteridium aquilinum var pseudocaudatum nymap.svg
NYFA-Z
USDA-N0
NEW 
ITIS
FNA Trop.


Images, wsp

Family Pteridaceae[edit]

The Pteridaceae E.D.M.Kirchn. (1831), (brakefern family)...

Subfamily Vittarioideae[edit]

Pteridaceaesubfam. Vittarioideae
Adiantum Maidenhair-fern N.Y. Status Images Distribution 1
 L.

1753. Adiantum pedatum  L.
Northern maidenhair-fern,
Northern maidenhair,
Five-finger fern

Adiante du Canada
Native,
Likely secure,
DEC-D[1]
Adiantum pedatum 3.jpg
Adiantum pedatum nymap.svg
NYFA-4
USDA-NN
NEW Can
ARS 
FNA Trop.


Images, wsp
Vittaria Shoestring-fern N.Y. Status Images Distribution 1
Farrar & Mickel (1991)
Appalachian gametophyte fern,
Appalachian shoestring fern,
Appalachian vittaria
Native,
Endangered,
DEC-A[2][3]
Vittaria appalachiana patch.JPG
Vittaria appalachiana nymap.svg
NYFA-1
USDA-N0





Images, wsp

Subfamily Cheilanthoideae[edit]

Pteridaceaesubfam. Cheilanthoideae
Pellaea Cliff-brake N.Y. Status Images Distribution 2
 (L.) Link

Pteris atropurpurea
Purple cliffbrake,
Purple-stemmed cliff-brake
Native,
Vulnerable,
DEC-D[1]
Pellaea atropurpurea concrete.jpg
Pellaea atropurpurea NY-dist-map.png
NYFA-4
USDA-NN





Images, wsp
 Mett. ex Kuhn
ssp. glabella

P. atropurpurea var. bushii
Smooth cliff-brake Native,
Threatened,
DEC-B[2]
Pellaea glabella fronds.jpg
Pellaea glabella ssp glabella NY-dist-map.png
NYFA-2
USDA-NN





Images, wsp
Cheilanthes Lip-fern N.Y. Status Images Distribution 1
(Michx.) D.C.Eaton

C. vestita (Spreng.) Sw.
Hairy lipfern,
Wooly lip-fern
Native,
Endangered,
DEC-A[3]
Cheilanthes lanosa 3 (FM).JPG
Cheilanthes lanosa NY-dist-map.png
NYFA-Z
USDA-N0





Images, wsp

Subfamily Cryptogrammoideae[edit]

Pteridaceaesubfam. Cryptogrammoideae
Cryptogramma Rock-brake N.Y. Status Images Distribution 1
(S.G.Gmel.) Prantl

1768. Pteris stelleri  S.G.Gmel.
1882. Cryptogramma stelleri  Prantl
Fragile rockbrake,
Slender cliff brake,
Slender rock-brake,
Steller's rock brake

Cryptogramme de steller
Native,
Vulnerable,
DEC-D[1]
Cryptogramma stelleri fronds.jpg
Cryptogramma stelleri NY-dist-map.png
NYFA-3-4
USDA-N
NEW Can
ARS ITIS
FNA Trop.
NSE 
IPN
Images, wsp

Subfamily Pteridoideae[edit]

Pteridaceaesubfam. Pteridoideae
Pteris Brake-fern N.Y. Status Images Distribution 1
 Poir.

1804. Pteris multifida  Poir.
1938. Pycnodoria multifida  (Poir.) Small
 auct.. Pteris serrulata non L.f.
Chinese brake,
Huguenot fern,
Saw-leaved bracken,
Spider brake,
Spider fern
Introduced from
 temperate Asia,
 tropical Asia,
No specimens
Pteris multifida inomotosou.JPG
Pteris multifida NY-dist-map.png
NYFA-X
USDA-X0

ARS ITIS
FNA Trop.

IPN
Images, wsp

Family Cystopteridaceae[edit]

The Cystopteridaceae Schmakov (2001) (bladder-fern family)...

Genus Cystopteris[edit]

Cystopteridaceaegenus Cystopteris
Cystopteris Bladder-fern N.Y. Status Images Distribution 5
 (L.) Bernh.

1753. Polypodium bulbiferum  L.
1806. Cystopteris bulbifera  (L.) Bernh.
1900. Filix bulbifera  (L.) Underw.
Bulblet bladder-fern,
Bulblet fern

Cystoptère bulbifère
Native,
Secure,
DEC-D[1]
Cystopterisbulbiferabulblet.jpg
Cystopteris bulbifera NY-dist-map.png
NYFA-5
USDA-NN





Images, wsp
 (L.) Bernh.

1753. Polypodium fragile  L.
1806. Cystopteris fragilis  (L.) Bernh.
1848. Cystopteris dickieana  R.Sim
1900. Filix fragilis  (L.) Underw.
Brittle bladder-fern,
Fragile fern
Native,
Likely secure,
DEC-D[1]
Cystopteris fragilis kz1.jpg
Cystopteris fragilis NY-dist-map.png
NYFA-4
USDA-NN





Images, wsp
 (Michx.) Desv.

Cystopteris protrusa ×
"Cystopteris hemifragilis"†

1803. Nephrodium tenue  Michx.
1827. Cystopteris tenuis  (Michx.) Desv.
1889. C. fragilis var. mackayi G.Lawson
Mackay's bladder-fern,
Mackay's fragile fern,
Upland brittle bladder-fern

Cystoptère ténue
Native,
Likely secure,
DEC-D[1]
Cystopteris tenuis-habitat.jpg
Cystopteris tenuis NY-dist-map.png
NYFA-4
USDA-NN





Images, wsp
 (Weath.) Blasdell

1935. C. fragilis var. protrusa Weath.
1963. Cystopteris protrusa  Blasdell
Lowland bladder-fern,
Lowland brittle fern,
Lowland fragile fern,
Southern bladder-fern
Native,
Endangered,
DEC-A[2]
Cystopteris protrusa-fronds.gif
Cystopteris protrusa NY-dist-map.png
NYFA-1
USDA-NN





Images, wsp
Cystopteris bulbifera × fragilis

Cystopteris bulbifera ×
Cystopteris fragilis
Hybrid fragile fern Native,
Unranked
Cystopteris bulbifera × fragilis nymap.svg
NYFA-U





Images, wsp

Genus Gymnocarpium[edit]

Cystopteridaceaegenus Gymnocarpium
Gymnocarpium Oak-fern N.Y. Status Images Distribution 1
 (L.) Newman

1753. Polypodium dryopteris  L.
1851. Gymnocarpium dryopteris  (L.) Newman
1852. Phegopteris dryopteris  (L.) Fée
1905. Dryopteris linnaeana  C. Chr.
1909. Dryopteris dryopteris  (L.) Christ
1917. Thelypteris dryopteris  (L.) Sloss.
1966. Carpogymnia dryopteris  Á.& D.Löve
Northern oak fern,
Common oak fern,
Western oak fern

Gymnocarpe
 fougère-du-chêne
Native,
Likely secure,
DEC-D[1]
Gymnocarpium dryopteris 6028 4x3.jpg
Gymnocarpium dryopteris nymap.svg
NYFA-4-5
USDA-NN
NEW 
ARS ITIS
FNA 


Images, wsp
(Sarvela) Pryer & Haufler

Gymnocarpium disjunctum ×
Gymnocarpium dryopteris

1980. G. dryopteris nssp. brittonianum  Sarvela
1993. G. × brittonianum  Pryer & Haufler
Britton's oak-fern Native,
Unranked
Gymnocarpium × brittonianum nymap.svg

USDA-NN

ITIS
FNA 


Images, wsp

Family Aspleniaceae[edit]

Genus Asplenium[edit]

The Aspleniaceae Newman (1849) (spleenwort family)...

Aspleniaceaegenus Asplenium
Asplenium Spleenwort N.Y. Status Images Distribution 10
var. platyneuron
Ebony spleenwort Native,
Secure,
DEC-D[1]
Asplenium platyneuron fronds.jpg
Asplenium platyneuron var platyneuron NY-dist-map.png
NYFA-5
USDA-NN
NEW 




Images, wsp
 L.

1753. Asplenium rhizophyllum  L.
1833. Camptosorus rhizophyllus  (L.) Link
Walking fern,
Walking-fern spleenwort,
American walking fern

Doradille ambulante,
Camptosore à
 feuilles radicantes
Native,
Likely secure,
DEC-D[1]
Asplenium rhizophyllum.jpg
Asplenium rhizophyllum NY-dist-map.png
NYFA-4-5
USDA-NN
NEW Can
ARS ITIS



Images, wsp
 L.
ssp. trichomanes
Maidenhair spleenwort Native,
Likely secure,
DEC-D[1]
Asplenium trichomanes - Capillaire - 1.jpg
Asplenium trichomanes ssp trichomanes NY-dist-map.png
NYFA-4
USDA-NN
NEW 




Images, wsp
 L.
ssp. quadrivalens D.E.Mey.
Maidenhair spleenwort Native,
Likely secure,
DEC-D[1]
Asplenium trichomanes muret Canejan (1).jpg
Asplenium trichomanes ssp quadrivalens NY-dist-map.png
NYFA-4
USDA-NN
NEW 




Images, wsp
 L.
nssp. lusaticum (D.E.Mey.) Lawalrèe

A. t. ssp. trichomanes ×
A. t. ssp. quadrivalens

1958. Asplenium × lusaticum  D.E.Mey.
1978. A. trichomanes nssp lusaticum
Hybrid
 Maidenhair spleenwort
Native,
Unranked,
No NY reports
Asplenium trichomanes muret Canejan.jpg
Nymap.svg

NEW 

Trop.


Images, wsp
 L.

1753. Asplenium ruta muraria  L.
1928. Asplenium cryptolepis  Fernald
1928. A. cryptolepis var. ohionis Fernald
1935. A. ruta-muraria var. cryptolepis
1939. A. ruta-muraria ssp. cryptolepis
Wallrue,
Wall-rue spleenwort

Doradille des murailles,
Mauerraute
Native,
Likely secure,
DEC-D[1]
Wall Rue, Gidcottmill Bridge - geograph.org.uk - 514113.jpg
Asplenium ruta-muraria nymap.svg
NYFA-4
USDA-NN
NEW Can
ARS ITIS
FNA Trop.


Images, wsp
Mountain spleenwort Native,
Threatened,
DEC-B[2]
BB-0066 Asplenium montanum.png
Asplenium montanum nymap.svg
NYFA-2-3
USDA-N0
NEW 




Images, wsp
 L.
var. americanum
(Fernald) Kartesz & Gandhi

1793. Scolopendrium vulgare  Small
1935. Phyllitis scolopendrium
   var.
americana Fernald
1954. Phyllitis fernaldiana  Á. Löve
1973. Phyllitis japonica ssp. americana
1991. A. scolopendrium var. americanum
American
 hart's-tongue fern,
Hart's-tongue fern,
Hart's tonguefern
Native,
Threatened,
DEC-B[2]
Asplenium scolopendrium americanum CR1934.jpg
Asplenium scolopendrium var americanum nymap.svg
NYFA-2
USDA-N

ARS 
FNA Trop.


Images, wsp
 L.
var. scolopendrium

1753. Asplenium scolopendrium  L.
1844. Phyllitis scolopendrium  Newman
1881. Scolopendrium
   scolopendrium
 H.Karst.
Hart's-tongue fern Introduced from
 Eurasia, n. Africa,
No specimens,
Cultivated
A fine Hart's-tongue Fern, Asplenium scolopendrium - geograph.org.uk - 1439635.jpg
Asplenium scolopendrium var scolopendrium nymap.svg


ARS 
Trop.


Images, wsp
 Huds.

1753. Asplenium ramosum  L.
1753. A. trichomanes-ramosum  L.
1762. Asplenium viride  Huds.
Green spleenwort,
Bright-green spleenwort
Native,
Endangered,
DEC-A[3]
Asplenium-viride.jpg
Asplenium viride nymap.svg
NYFA-1
USDA-NN
NEW 

Trop.


Images, wsp
 D.C.Eaton

Asplenium montanum ×
Asplenium platyneuron

1873. Asplenium bradleyi  D.C.Eaton
1925. Asplenium stotleri  Wherry
1931. Chamaefilix bradleyi  Farw.
Bradley's spleenwort Native,
No recent reports,
Endangered,
DEC-A[3]
BB-0068 Asplenium bradleyi.png
Asplenium bradleyi nymap.svg
NYFA-Z
USDA-N0

ITIS
FNA 


Images, wsp
 R.R.Scott

Asplenium rhizophyllum ×
Asplenium platyneuron

1865. Asplenium ebenoides  R.R.Scott
1937. × Asplenosorus ebenoides  Wherry
Scott's spleenwort,
Dragon tail fern,
Walking spleenwort
Native,
No recent reports
BB-0058 Asplenium ebenoides.png
Asplenium × ebenoides nymap.svg
NYFA-Z
USDA-N0
NEW 




Images, wsp
 Syme

Asplenium ruta-muraria ×
Asplenium trichomanes

1886. Asplenium clermontae  Syme
Hybrid
 Wallrue &
 Maidenhair spleenwort
Native,
Unranked,
No NY reports
Asplenium ×‌ clermontae nymap.svg
NYFA-D
USDA-N0
NEW 




Images, wsp

Family Thelypteridaceae[edit]

The Thelypteridaceae Pic.Serm (1970) (marsh fern family)

Thelypteridaceae
Phegopteris Beech-fern N.Y. Status Images Distribution 2

Dryopteris phegopteris
Lastrea phegopteris
Phegopteris polypodioides
Polypodium connectile
Thelypteris phegopteris
Northern beech fern Native,
Secure,
DEC-D[1]
Phegopteris connectilis 2.jpg
Phegopteris connectilis NY-dist-map.png
NYFA-5





Images, wsp

Dryopteris hexagonoptera
Polypodium hexagonopterum
Thelypteris hexagonoptera
Broad beech fern Native,
Likely secure,
DEC-D[1]
Phegopteris hexagonoptera1.JPG
Phegopteris hexagonoptera NY-dist-map.png
NYFA-4-5





Images, wsp
Thelypteris Maiden-fern N.Y. Status Images Distribution 3

Dryopteris noveboracensis
Polypodium noveboracense
Thelypteris thelypterioides
New York fern Native,
Secure,
DEC-D[1]
Thelypteris noveboracensis.jpg
Thelypteris noveboracensis NY-dist-map.png
NYFA-5





Images, wsp
var. pubescens

Dryopteris thelypteris
Lastrea thelypteris
Thelypteris confluens
  var. pubescens
Marsh fern Native,
Secure,
DEC-D[1]
Thelypteris palustris Russia 3.jpg
Thelypteris palustris var pubescens NY-dist-map.png
NYFA-5





Images, wsp

Aspidium simulatum
Dryopteris simulata
Massachusetts fern,
Bog fern
Native,
Vulnerable,
DEC-D[1]
BB-0039 Dryopteris simulata.png
Thelypteris simulata NY-dist-map.png
NYFA-3-4
USDA-N





Images, wsp

Family Woodsiaceae[edit]

The Woodsiaceae Herter (1949) (cliff fern family)...

Woodsiaceae
Woodsia Cliff-fern N.Y. Status Images Distribution 5
 (L.) R.Br.

Acrostichum ilvense L
Rusty cliff fern,
Rusty woodsia,
woodsie de l'île d'Elbe
Native,
Secure,
DEC-D[1]
Woodsia ilvensis on serpentine.JPG
Woodsia ilvensis NY-dist-map.png
NYFA-5
USDA-NN





Images, wsp
 (Spreng.) Torrey
ssp. obtusa
Bluntlobe cliff fern,
Blunt-lobe woodsia
Native,
Secure,
DEC-D[1]
Woodsia obtusa blade.jpg
Woodsia obtusa ssp obtusa nymap.svg
NYFA-5
USDA-NN





Images, wsp
 (Bolt.) S.Gray

Acrostichum alpinum
W. alpina var. bellii
W. glabella var. bellii
Alpine cliff fern,
Alpine woodsia,
Northern woodsia

Woodsie alpine
Native,
Endangered,
DEC-A[2]
BB-0024 Woodsia alpina.pngImg spacer 2x87.pngWoodsia hyperborea Atlas Alpenflora.jpg
Woodsia alpina nymap.svg
NYFA-1
USDA-NN

ITIS



Images, wsp
R.Br. ex Richardson (1823)

W. alpina var. glabella
W. hyperborea var. glabella
Smooth cliff fern,
Smooth woodsia

Woodsie glabre
Native,
Endangered,
DEC-A[2]
BB-0025 Woodsia glabella.png
Woodsia glabella nymap.svg
NYFA-1
USDA-NN


FNA 


Images, wsp
 D.C.Eaton (1865)
ssp. cathcartiana

W. cathcartiana B.L.Rob. (1908)
W. oregana var. cathcartiana
Oregon cliff fern,
Oregon woodsia,
(tetraploid)
Native,
Likely extirpated
Woodsia oregana.jpg
Woodsia oregana ssp cathcartiana nymap.svg
NYFA-Z
USDA-NN

ITIS



Images, wsp

Family Onocleaceae[edit]

The Onocleaceae ...

Onocleaceae
Onoclea Sensitive fern N.Y. Status Images Distribution 1
 L. (1753)
Sensitive fern,
Bead fern,
Onoclée sensible
Native,
Secure
Onoclea sensibilis.jpg
Onoclea sensibilis nymap.svg
NYFA-5
USDA-NN

ARS ITIS
FNA 


Images, wsp
Matteuccia Ostrich fern N.Y. Status Images Distribution 1
 (L.) Tod.
var. pensylvanica (Willd.) C.V.Morton

Matteuccia pensylvanica
Struthiopteris pensylvanica
Pteretis pensylvanica
Onoclea struthiopteris pensylvanica
Ostrich fern,
Shuttlecock fern,
Fiddlehead fern

Fougère à l'autruche
Native,
Likely secure
Jonpapartis001.JPG
Matteuccia struthiopteris var pensylvanica nymap.svg
NYFA-4
USDA-NN
NEW 
ARS ITIS
FNA 


Images, wsp

Family Blechnaceae[edit]

The Blechnaceae Newman (1844) ...

Blechnaceae
Woodwardia Chainfern N.Y. Status Images Distribution 2
Virginia chainfern Native,
Secure,
DEC-D[1]
Woodwardia virginica.jpgImg spacer 2x87.pngWovi 004 lvp.jpg
Woodwardia virginica NY-dist-map.png
NYFA-5
USDA-N





Images, wsp
Netted chainfern Native,
Vulnerable,
DEC-D[1]
Woodwardia areolata.jpeg
Woodwardia areolata nymap.svg
NYFA-3
USDA-N





Images, wsp

Family Athyriaceae[edit]

The Athyriaceae Alston (1956) (lady-fern family)...

Athyriaceae
Deparia False-spleenwort N.Y. Status Images Distribution 1
 (Sw.) M.Kato

Athyrium thelypterioides
Diplazium acrostichoides
Silvery spleenwort Native,
Secure,
DEC-D[1]
BB-0069 Athyrium thelypteroides.pngImg spacer 2x87.pngGjk-deparia thelypterioides.gif
Deparia acrostichoides nymap.svg
NYFA-5
USDA-NN





Images, wsp
DiplaziumSw. Twin-sorus fern N.Y. Status Images Distribution 1
 (Spreng.) Broun

1804. Asplenium pycnocarpon  Spreng.
1906. Athyrium pycnocarpon  Tidestr.
1938. Diplazium pycnocarpon  M.Broun
1977. Homalosorus pycnocarpos  Serm.
Glade fern,
Narrow-leaved glade fern,
Narrow-leaved-spleenwort

Athyrie à sores denses,
Diplazie à sores denses
Native,
Likely secure,
DEC-D[1]
Diplazium pycnocarpon 1924.jpgImg spacer 2x87.pngDiplazium pycnocarpon 1961.jpg
Diplazium pycnocarpon nymap.svg
NYFA-4
USDA-NN
NEW Can
ITIS
FNA Trop.


Images, wsp
AthyriumRoth Lady-fern N.Y. Status Images Distribution 2
 (L.) Roth ex Mert.
ssp. angustum (Willd.) R.T.Clausen

1810. Aspidium angustum  Willd.
1825. Athyrium angustum  (Willd.) C.Presl
1864. A. filix-femina var. angustum G.Lawson
1949. A. filix-femina ssp. angustum R.T.Clausen
Common lady fern,
Narrow lady fern,
Northern lady fern,
Subarctic lady fern
Native,
Secure,
DEC-D[1]
Athyrium filix -femina.jpg
Athyrium filix-femina ssp angustum nymap.svg
NYFA-5
USDA-NN
NEW Can
ARS ITIS
FNA Trop.
NSE 
IPN
Images, wsp
 (L.) Roth ex Mert.
ssp. asplenioides (Michx.) Hultén

1803. Nephrodium asplenioides  Michx.
1817. Athyrium asplenioides  (Michx.) A.A.Eaton.
1923. A. filix-femina var. asplenioides Farw,
1962. A. filix-femina ssp. asplenioides Hultén
Southern lady fern,
Lowland lady fern
Native,
Vulnerable,
DEC-D[1]
XN Athyrium filix-femina.jpg
Athyrium filix-femina ssp asplenioides nymap.svg
NYFA-3
USDA-N0
NEW 
ITIS
FNA Trop.
NSE 

Images, wsp
 (L.) Roth ex Mert.
ssp. angustum × asplenioides

A. filix-femina ssp. angustum ×
A. filix-femina ssp. asplenioides
Hybrid
 Common lady fern &
 Southern lady fern
Native,
Unranked,
No sepecimens
Athyrium angustum.jpgImg spacer 2x87.pngAthyrium filix-femina (Leaf).jpg
Athyrium filix-femina ssp angustum × ssp asplenioides nymap.svg
NYFA-D





Images, wsp
AnisocampiumC.Presl An jue shu N.Y. Status Images Distribution 1
(Mett.) Yea C.Liu, W.L.Chiou & M.Kato

1865. Asplenium nipponicum  Mett.
1873. Athyrium nipponicum  (Mett.) Hance
2011. Anisocampium niponicum [2]
Japanese painted lady fern,
Japanese painted fern
Introduced from
 e. Asia,
No specimens
Athyrium niponicum f metallicum1.jpg
Anisocampium niponicum nymap.svg
NYFA-0
NEW 
ARS 
Trop.


Images, wsp
  1. a b c d New York State Department of Environmental Conservation (DEC) Regulation §193.3, Protected Native Plants, (d) Exploitably vulnerable.
  2. [ Liu & al. (2011) "Molecular phylogeny and taxonomy of the fern genus Anisocampium (Athyriaceae)". TAXON 60 (3), June 2011: 824–830.]

Family Dryopteridaceae[edit]

The Dryopteridaceae Herter (1949) (woodfern family)...

Genus Cyrtomium[edit]

Dryopteridaceaegenus Cyrtomium
Cyrtomium Netvein-hollyfern N.Y. Status Images Distribution 1

Polypodium falcatum
Japanese holly fern,
Japanese netvein hollyfern
Introduced
Cyrtomium falcatum 001.jpg
Cyrtomium falcatum nymap.svg
NYFA-X
USDA-X





Images, wsp

Genus Dryopteris[edit]

North American Dryopteris reticulogram.svg

Most Dryopteris (wood fern) species are characterized by sori located near the mid leaf vein. Dryopteris hybrids are very common.

Dryopteris diploid species[edit]

These species have 2n=82 chromosmes (twice the Dryopteris base chromosome number of n=41). D. intermedia and D. Goldiana have been shown to be part of a North American "reticulate complex", in which, along with three other Dryopteris species, they are parents to most of the allopolyploid Dryopteris species shown below.[1][2]

Dryopteridaceaegenus Dryopteris
Dryopteris Woodfern N.Y. Status Images Distribution 30
 (L.) A.Gray

1753. Polypodium marginale  L.
1848. Dryopteris marginalis  A.Gray
Marginal wood fern Native,
Secure,
DEC-D[3]
DryopterisMarginalis.jpg
Dryopteris marginalis NY-dist-map.png
NYFA-5
USDA-NN
NEW 

FNA 


Images, wsp
(Muhl. ex Willd.) A. Gray
ssp. intermedia

D. spinulosa var. intermedia
Evergreen woodfern Native,
Secure,
DEC-D[3]
BB-0048 Dryopteris intermedia.pngImg spacer 2x87.pngFig 2 Dryopteris spinulosa intermedia.png
Dryopteris intermedia ssp intermedia NY-dist-map.png
NYFA-5
USDA-NN
NEW 

FNA 


Images, wsp
(Hook. ex Goldie) A.Gray

1822. Aspidium goldieanum  Hook. ex Goldie
1848. Dryopteris goldieana  A.Gray
1910. Thelypteris goldieana  Nieuwl.
1916. Filix goldiana  Farw.
1931. Filix-mas goldiana  Farw.
Goldie's woodfern,
Goldie's shield fern,
Giant woodfern

Dryoptère de Goldie,
Dryoptéride de Goldie,
Dryoptéris géant
Native,
Likely secure,
DEC-D[3]
BB-0043 Dryopteris goldiana.png
Dryopteris goldiana NY-dist-map.png
NYFA-4
USDA-NN
NEW Can
ARS ITIS
FNA Trop.
NSE 
IPN
Images, wsp
 (L.) Schott
Fragrant cliff wood-fern,
Fragrant woodfern
Native,
Endangered,
DEC-A[4]
Dryopteris fragrans 1.jpg
Dryopteris fragrans nymap.svg
NYFA-1
USDA-NN
NEW 

FNA 


Images, wsp
Dryopteris allopolyploid species[edit]

These are fertile species of hybrid origin. Most are allotetraploid, derived from diploid Dryopteris parents, including two of the New York diploid species, D. intermedia and D. goldiana, shown above. The other diploid parents, D. expansa and D. ludoviciana, are North American, but not New York natives. A fifth, likely extinct, diploid parent "D. semicristata" has been hypothesized to form the well-supported model used here. The species Dryopteris clintoniana is allohexaploid, resulting from the diploid D. goldiana (above) and the allotetraploid D. Cristata, below. The parentage of the introduced non-NY-native, Dryopteris filix-mas has not been resolved, but it is probably not part of the same North American "reticulate complex" as the rest.[1]

Dryopteridaceaegenus Dryopteris
Dryopteris Woodfern N.Y. Status Images Distribution 30
 (L.) A.Gray

♀ D. "semicristata" (SS) ×
♂ D. ludoviciana (LL) → SSLL

1753. Polypodium cristatum  L.
1848. Dryopteris cristata  A.Gray
Crested shield-fern,
Crested woodfern
Native,
Secure,
DEC-D[2]
Farnwedel 165a.jpgImg spacer 2x87.pngBB-0041 Dryopteris cristata.png
Dryopteris cristata NY-dist-map.png
NYFA-5
USDA-NN
NEW 

FNA 


Images, wsp
(Vill.) H.P.Fuchs

♀ D. "semicristata" (SS) ×
♂ D. intermedia (II) → SSII

1777. Polypodium spinulosum  O.F.Muell.
1786. Polypodium carthusianum  Vill.
1867. Dryopteris spinulosa  Watt
1959. Dryopteris carthusiana  H.P.Fuchs
Spinulose shield fern,
Spinulose wood fern,
Toothed wood fern,
Narrow buckler fern

Dryoptère spinuleuse
Native,
Secure,
DEC-D[2]
Dryopteris carthusiana3.jpg
Dryopteris carthusiana NY-dist-map.png
NYFA-5
USDA-N
NEW Can
ARS 
FNA 


Images, wsp
(D.C.Eaton) Dowell

♀ D. cristata (SSLL) ×
♂ D. goldiana (GG) → SSLLGG

1867. Aspidium cristatum
   var.
clintonianum D.C.Eaton
1893. Dryopteris cristata
   var.
clintoniana Underw.
1906. Dryopteris clintoniana  Dowell
1920. Thelypteris clintoniana  House
Clinton's wood fern

Dryoptère de clinton
Native,
Secure,
DEC-D[2]
Dryopteris clintoniana cultivated.JPG
Dryopteris clintoniana NY-dist-map.png
NYFA-5
USDA-NN
NEW 

FNA Trop.


Images, wsp
(Kunze) Clarkson

♀ D. intermedia (II) ×
♂ D. expansa (EE) → IIEE

1848. Aspidium campylopterum  Kunze
1915. Dryopteris austriaca  Woyn.
1915. D. spinulosa var. americana Fernald
1916. D. dilatata var. americana Benedict
1920. Thelypteris dilatata  (Hoffm.) House
1930. D. campyloptera  Clarkson
Mountain woodfern,
Spreading wood fern,
Eastern spreading wood fern

Dryoptère arquée
Native,
Likely secure,
DEC-D[2]
Dryopteris campyloptera frond.JPG
Dryopteris campyloptera NY-dist-map.png
NYFA-4
USDA-N
NEW Can
ARS ITIS
FNA Trop.


Images, wsp
 (W.Palmer)
Knowlt., Palmer & Pollard ex Small

♀ D. ludoviciana (LL) ×
♂ D. goldiana (GG) → LLGG

1899. D. goldiana ssp. celsa W.Palmer
1900. Dryopteris celsa  Knowlt. &al.
1938. Dryopteris atropalustris  Small
1960. Dryopteris wherryi  Crane
Log fern Native,
Endangered,
DEC-A[3]
Dryopteris celsa, from Log Fern 2356c.jpg
Dryopteris celsa nymap.svg
NYFA-1
USDA-N

ARS ITIS
FNA Trop.


Images, wsp
 (L.) Schott

1753. Polypodium filix-mas  L.
1801. Nephrodium filix-mas  Rich.
1834. Dryopteris filix-mas  Schott
Male fern,
Male wood-fern
Introduced,
N. America native
Dryopteris filix-mas 001.jpg
Dryopteris filix-mas nymap.svg
NYFA-X
USDA-NN
NEW 
ARS 
FNA Trop.
NSE BNA 

Images, wsp
Dryopteris named hybrids[edit]
Dryopteridaceaegenus Dryopterissubsect. named hybrids
Dryopteris Woodfern N.Y. Status Images Distribution 30
Dryopteris × boottii
 (Tuck.) Underw.

D. cristata (SSLL) ×
D. intermedia (II) → ILS

1843. Aspidium boottii  Tuck.
1893. Dryopteris boottii  (Tuck.) Underw.
1910. Thelypteris boottii  (Tuck.) Nieuwl.
Boott's wood fern,
Glandular swamp fern
Native,
Vulnerable
Dryopteris × boottii BB-1913.png
Dryopteris × boottii NY-dist-map.png
NYFA-3-5
USDA-NN
NEW 
ITIS



Images, wsp
Dryopteris × triploidea
 Wherry

D. carthusiana (IISS) ×
D. intermedia (II) → IIS

1930. D. spinulosa var. fructuosa Trudell
1950. D. austriaca var. fructuosa C.V.Morton
1958. D. intermedia var. fructuosa Wherry
1960. Dryopteris × triploidea  Wherry
Triploid woodfern Native,
Vulnerable
BB-0046 Dryopteris spinulosa.pngImg spacer 2x87.pngBB-0048 Dryopteris intermedia.png
Dryopteris × triploidea NY-dist-map.png
NYFA-3
USDA-NN

ITIS
FNA 


Images, wsp
Dryopteris × uliginosa
(A.Braun ex Dowell) Druce

D. carthusiana (IISS) ×
D. cristata (SSLL) → ILSS
Spinulose crested wood fern Native,
Vulnerable
BB-0046 Dryopteris spinulosa.pngImg spacer 2x87.pngBB-0041 Dryopteris cristata.png
Dryopteris × uliginosa NY-dist-map.png
NYFA-3
USDA-NN





Images, wsp
Dryopteris × slossoniae
Wherry ex Lellinger

D. cristata (SSLL) ×
D. marginalis (MM) → LMS

1894. D. cristata × marginalis  Davenport
1894. Aspidium cristatum × marginale
1904. Nephrodium slossonae  Hahne
1942. Dryopteris × slossonae  Wherry
Crested marginal fern,
Miss Slosson's wood fern
Native,
Threatened
BB-0041 Dryopteris cristata.pngImg spacer 2x87.pngBB-0044 Dryopteris marginalis.png
Dryopteris × slossoniae NY-dist-map.png
NYFA-1-3
USDA-NN





Images, wsp
Dryopteris × pittsfordensis
 Slossen

D. carthusiana (IISS) ×
D. marginalis (MM) → IMS

1904. Dryopteris pittsfordensis  Sloss.
1904. Nephrodium pittsfordensis  Woolson
Pittsford's wood fern Native,
Endangered
Fig 1 Dryopteris pittsfordensis.png
Dryopteris × pittsfordensis nymap.svg
NYFA-1
USDA-NN

ITIS
Trop.

IPN
Images, wsp
Dryopteris × burgessii
 Boivin

D. clintoniana (SSLLGG) ×
D. marginalis (MM) → GLMS
Burgess' shield fern Native,
No recent reports
BB-0042 Dryopteris clintoniana.pngImg spacer 2x87.pngBB-0044 Dryopteris marginalis.png
Dryopteris × burgessii NY-dist-map.png
NYFA-Z?
USDA-NN





Images, wsp
Dryopteris × benedictii
 (Farw.) Wherry

D. carthusiana (IISS) ×
D. clintoniana (SSLLGG) → GILSS
Benedict's wood fern Native,
No recent reports
BB-0046 Dryopteris spinulosa.pngImg spacer 2x87.pngBB-0042 Dryopteris clintoniana.png
Dryopteris × benedictii NY-dist-map.png
NYFA-Z?
USDA-NN





Images, wsp
Dryopteris × dowellii
 (Farw.) Wherry

D. clintoniana (SSLLGG) ×
D. intermedia (II) → GILS
Dowell's wood fern Native,
No recent reports
BB-0042 Dryopteris clintoniana.pngImg spacer 2x87.pngBB-0048 Dryopteris intermedia.png
Dryopteris × dowellii NY-dist-map.png
NYFA-Z
USDA-NN





Images, wsp
Dryopteris × mickelii

D. clintoniana (SSLLGG) ×
D. goldiana (GG) → GGLS
Wood fern Native,
Likely extirpated
BB-0042 Dryopteris clintoniana.pngImg spacer 2x87.pngBB-0043 Dryopteris goldiana.png
Dryopteris × mickelii NY-dist-map.png
NYFA-Z
USDA-N





Images, wsp
Dryopteris × neowherryi
 W.H.Wagner

D. goldiana (GG) ×
D. marginalis (MM) → GM
Wood fern Native,
No recent reports
BB-0043 Dryopteris goldiana.pngImg spacer 2x87.pngBB-0044 Dryopteris marginalis.png
Dryopteris × neowherryi NY-dist-map.png
NYFA-Z
USDA-NN





Images, wsp
Dryopteris × separabilis
 Small (pro sp.)

D. celsa (LLGG) ×
D. intermedia (II) → GIL
Glandular log fern Native,
No recent reports
BB-0048 Dryopteris intermedia.png
Dryopteris × separabilis NY-dist-map.png
NYFA-Z
USDA-N0





Images, wsp
Dryopteris × correllii
 W.H.Wagner

D. carthusiana (IISS) ×
D. goldiana (GG) → GIS
Correll's wood fern,
Poyser's wood fern
Native,
No recent reports
BB-0046 Dryopteris spinulosa.pngImg spacer 2x87.pngBB-0043 Dryopteris goldiana.png
Dryopteris × correllii nymap.svg
NYFA-?
USDA-N0





Images, wsp
Dryopteris × leedsii
 Wherry

D. celsa (LLGG) ×
D. marginalis (MM) → GLM
Leeds' wood fern Native,
No recent reports
BB-0043 Dryopteris goldiana.pngImg spacer 2x87.pngBB-0044 Dryopteris marginalis.png
Dryopteris × leedsii nymap.svg
NYFA-Z
USDA-N0
NEW 
ITIS



Images, wsp
Dryopteris unnamed hybrids[edit]
Dryopteridaceaegenus Dryopterissubsect. unnamed hybrids
Dryopteris Woodfern N.Y. Status Images Distribution 30
Dryopteris filix-mas × marginalis
Marginal male fern Native,
Threatened
BB-0045 Dryopteris filix-mas.pngImg spacer 2x87.pngBB-0044 Dryopteris marginalis.png
Dryopteris filix-mas × marginalis nymap.svg
NYFA-1-3





Images, wsp
Dryopteris celsa × goldiana

D. celsa (LLGG) ×
D. goldiana (GG) → GGL
Hybrid wood-fern Native,
No recent reports
BB-0043 Dryopteris goldiana.png
Dryopteris celsa × goldiana nymap.svg
NYFA-Z





Images, wsp
Dryopteris intermedia × marginalis

D. intermedia (II) ×
D. marginalis (MM) → IM
Glandular marginal fern Native,
No recent reports
BB-0048 Dryopteris intermedia.pngImg spacer 2x87.pngBB-0044 Dryopteris marginalis.png
Dryopteris intermedia × marginalis nymap.svg
NYFA-Z





Images, wsp
Dryopteris celsa × cristata

D. celsa (LLGG) ×
D. cristata (SSLL) → GLLS
Hybrid wood-fern Native,
No recent reports
BB-0041 Dryopteris cristata.png
Dryopteris celsa × cristata nymap.svg
NYFA-Z





Images, wsp
Dryopteris carthusiana × celsa

D. carthusiana (IISS) ×
D. celsa (LLGG) → IGLS
Spinulose wood fern Native,
No recent reports
BB-0046 Dryopteris spinulosa.png
Dryopteris carthusiana × celsa nymap.svg
NYFA-Z





Images, wsp
Dryopteris goldiana × intermedia

D. goldiana (GG) ×
D. intermedia (II) → GI
Wood fern Native,
No recent reports
BB-0043 Dryopteris goldiana.pngImg spacer 2x87.pngBB-0048 Dryopteris intermedia.png
Dryopteris goldiana × intermedia nymap.svg
NYFA-Z





Images, wsp
Dryopteris cristata × goldiana

D. cristata (SSLL) ×
D. goldiana (GG) → GLS
Benedict's wood fern Native,
Unranked
BB-0041 Dryopteris cristata.pngImg spacer 2x87.pngBB-0043 Dryopteris goldiana.png
Dryopteris cristata × goldiana nymap.svg
NYFA-U





Images, wsp

Genus Polystichum[edit]

Polystichum goes by the English vernacular names Chrismas fern, holy fern, and sword fern.

Dryopteridaceaegenus Polystichum
Polystichum Hollyfern N.Y. Status Images Distribution 4
 (Michx.) Schott
var. acrostichoides

1803. Nephrodium acrostichoides  Michx.
1834. Polystichum acrostichoides  Schott
Christmas fern,
Dagger fern

Polystic faux-acrostiche
Native,
Secure
Christmas Fern Polystichum acrostichoides Plant 2000px.JPGImg spacer 2x87.pngBB-0034 Polystichum acrostichoides.png
Polystichum acrostichoides var acrostichoides NY-dist-map.png
NYFA-5
USDA-NN

ARS 
Trop.


Images, wsp
 (Spenner) Fée

1825. Aspidium braunii  Spenner
1852. Polystichum braunii  Fée
Braun's holly-fern Native,
Likely secure
Polystichum braunii 1-OB8.jpg
Polystichum braunii NY-dist-map.png
NYFA-4
USDA-NN


Trop.


Images, wsp
 Barrington
[ P. acrostichoides × P. braunii ]

1986. Polystichum × potteri  Barrington
Hybrid Christmas fern
  & Braun's holly-fern
Native,
Unranked
Christmas Fern Polystichum acrostichoides Leaf 2000px.JPGImg spacer 2x87.pngBB-0036 Polystichum braunii.png
Polystichum × potteri nymap.svg
NYFA-U
USDA-NN


Trop.


Images, wsp
 (L.) Roth

1753. Polypodium lonchitis  L.
1800. Polystichum lonchitis  Roth
1801. Aspidium lonchitis  Sw.
1924. Polystichum mohrioides var. lemmonii
Northern holly-fern

Polystic faux-lonchitis
Native,
Endangered
Lanzen-Schildfarn Polystichum lonchitis Grüner Ups Ehrwald-003.jpg
Polystichum lonchitis nymap.svg
NYFA-Z
USDA-NN


FNA 


Images, wsp
 Cody
[ P. acrostichoides × P. lonchitis ]

1924. Polystichum lonchitis fo. marginale McColl
1966. Polystichum marginale  (McColl) Cody
1968. Polystichum × hagenahii  Cody
Hagenah's holly fern,
Hagenah's polystichum,
Hybrid Christmas fern
  & northern holly-fern

Polystic de Hagenah
Native,
Unranked,
Questionable[1]
Christmas Fern Polystichum acrostichoides Unfurled 2000px.jpgImg spacer 2x87.pngPolystichum lonchitis 5576.JPG
Polystichum × hagenahii nymap.svg


ITIS
FNA 


Images, wsp
  1. Flora of North America states that the rare hybrid Polystichum × hagenahii has only been found growing with both parents in Ontario. NYFA does not list it, but USDA-PLANTS does show it in NY as well as PA, listing the NY source: Lellinger, D.B. (1985) A field manual of the ferns and fern-allies of the U.S. and Canada, Smithsonian Institution, Washington, DC.

Family Polypodiaceae[edit]

The Polypodiaceae J.Presl & C.Presl (1822) (polypody family)

Polypodiaceae
Polypodium Polypody N.Y. Status Images Distribution 3
 L.

1753. Polypodium virginianum  L.
1818. P. vulgare var. virginianum Eaton
1840. P. vulgare var. americanum Hook.
1962. P. vulgare ssp. virginianum Hultén
1977. P. vinlandicum  Á. Löve & D. Löve
Rock polypody,
Rock-cap fern,
Common polypody

Tripes-de-roches,
Polypode de virginie
Native,
Secure
Polypodium virginianum3.jpg
Polypodium virginianum NY-dist-map.png
NYFA-5
USDA-NN
NEW 

FNA 


Images, wsp
Haufler & Windham

1900. P. vulgare var. oreophilum Maxon
1902. P. vulgare var. acuminatum Gilbert
1922. P. virginianum fo. acuminatum Fernald
1991. P. appalachianum  Haufler & Windham
Appalachian rockcap fern,
Appalachian polypody,
Appalachian rock polypody

Polypode des Appalaches
Native,
Secure
Polypodium appalachianum1.jpg
Polypodium appalachianum NY-dist-map.png
NYFA-5
USDA-NN
NEW 
ARS ITIS
FNA Trop.


Images, wsp
 Cusick
[ P. appalachianum × P. virginianum ]

2002. Polypodium × incognitum  Cusick
Hybrid polypody Native,
Unranked
Polypodium virginianum (5382723883).jpg
Polypodium × incognitum nymap.svg
NYFA-?
NEW 




Images, wsp
Pleopeltis Scaly-polypody N.Y. Status Images Distribution 1
(L.) Andrews & Windham
var. michauxiana
(Weath.) Andrews & Windham

1939. Polypodium polypodioides
   var.
michauxianum Weath.
1993. Pleopeltis polypodioides
   var.
michauxiana Andrews & Windham
Resurrection fern,
Gray's polypody
Introduced,
N. America native,
Questionable[1]
Pleopeltis polypodioides Brooker Creek.jpg
Pleopeltis polypodioides var michauxiana nymap.svg
NYFA-X
USDA-N0


FNA 


Images, wsp
  1. USDA-NRCS-PLANTS includes New York in the range of Pleopeltis polypodioides, but the reference for this (House, H.D. 1924. Annotated list of the ferns and flowering plants of New York State. New York State Museum Bulletin No. 254) mentions only that the "gray polypody, or tree fern (Polypodium polypodioides) (L.) Watt, has been found on driftwoud on the Staten Island shore, although its natural range is from Virginia southward."